Dedications

We dedicate the present work to Professor Simonas Pileckis, who laid the foundation of coleopterology in Lithuania and taught us to discover the wonders of the beetle’s world. I also dedicate this paper to my parents, Jurgiui and Janinai Tamučiams, for encouraging my childhood interests in natural history (VT).

Introduction

The Order Coleoptera, or beetles, forms the most numerous group of insects throughout the animal kingdom. In ecological terms this is a very diverse group of insects occupying a wide variety of terrestrial and freshwater ecological niches. The value of these animals to terrestrial ecosystems and to humankind is enormous, and thus, knowledge of fauna is of particular importance.

Faunistic research is essential in assessing biodiversity in a given area and also for understanding processes at present occurring in nature related to the increasing anthropogenic impact and climate change. Faunistic research data are important not only to fundamental sciences, but it also can be widely used in many fields of applied sciences such as agriculture, forestry, ecology, environmental protection and many others. Inventories of such data and making a checklist are the most important steps to make the data accessible and suitable for use by the general public. Moreover, faunistic studies have both scientific and cultural-social importance. The above-mentioned aspects directly show the awareness of society about own country’s natural history and the necessity of its conservation.

The last 20 years has been a time of major changes in beetle classification. It can be summarized from the reviewed literature that over 600 beetle species’ names have been changed since their publication in the previous monograph of Lithuanian beetles (Pileckis and Monsevičius 1995, revised 1997) due to reidentification and spelling emendations. Such situation causes increasing discrepancies in nomenclature of beetles species used by various authors in the Lithuanian faunistic literature. This in turn leads to numerous misunderstandings and is highly inconvenient for anybody working with faunal lists, especially one who is not well acquainted with current beetle literature. Moreover, the necessity to summarize all the available literature on Lithuanian beetles was felt for a long time. A lot of information scattered in various publications was forgotten and unusable in the studies of Lithuanian Coleoptera. Therefore, the publication of a new, updated checklist became an urgent necessity. To fill this gap we decided not only to compile the list of Lithuanian beetles, but also to summarize all available literature sources on Lithuanian beetle fauna, to create the bibliographic list, and to present reference sources for each species separately. In addition, we predicted the expected species for Lithuania based on their presence in faunas of neighboring republics and considering conditions in these countries from both geographic and ecological aspects. Analyzing beetle fauna of such large countries as Sweden, Poland and Belarus, we considered the importance of assessing the diversity of natural systems in large territories. Species predicted for Lithuania are found in regions adjacent to our country, where natural conditions are similar to those in Lithuania; therefore, there is a high likelihood of detection in Lithuania. We believe that these forecastsprovide directions to faunistic research and facilitate the search of new species. Additionaly, we decided to comment on the main changes in beetle classification as well as mention the untrustworthy notifications of some species listed in the recently published multivolume Catalogue of Palaearctic Coleoptera, Web project “Fauna Europaea Web Service, ” various regional catalogues and species lists. We hope this work will encourage researchers to look deeper into species distribution over the whole territory of the Republic. Considering distribution ranges of some species and their revised identifications, we suggest to remove 43 species from the list of Lithuanian beetles. We hope the list of beetles accomplished by us and presented in this paper will be useful for regional research in applied and conservation studies of beetles. Furthermore, it is our intention to help both Lithuanian and foreign entomologists to realize a truer picture of the current beetle diversity in Lithuania and to present a species list which is up to date as of 2010, with all doubtful entries indicated. We reference herein all new taxonomic placements and provide explanations for every nomenclatural change compared to those published in two volumes Lithuanian Fauna. The Beetles (Pileckis and Monsevičius 1995, 1997).

Unfortunately we failed to avoid some shortcomings in our paper. In our opinion one of the major disadvantages is that the information presented is based mostly only on published sources. Much of the faunistic data is accumulated in the database and collections of Kaunas T. Ivanauskas zoological museum, other collections stored in the Lithuanian universities or private collections, in the museums of foreign countries, but it is impossible physically for us to process the data and present it in this catalogue. In our view it will be much better to revise particular families or subfamilies and to use this data in the future and we sincerely hope that our catalogue will stimulate such process.

Characterisation of territory

The main Lithuanian geomorphological features formed during the several glacial periods which started in the Pleistocene Epoch. The last glacier covered almost all Lithuania except the southeastern part. During this period large moraine structures formed from glacial drifts (Česnulevičius 1998). Morainae glacial lakes and the old alluvial plains, morainal hills, littoral plains, zandric plains and river valleys dominate in the Lithuanian landscape. Sand dunes stretch in the western country which is in the Curonian Spit and the continental seaside. Hilly and lake-rich landscape dominates in eastern, southern and western hills. There are various areas of agriculture, forests, lakes and river valleys. Marshes, swamps and other wetlands are common in lowlands and valleys. Wide and the homogeneous moraine and limnoglacial plains landscape is characteristic in central, northern and southeastern Lithuania. In these regions agriculture is developed because of rich soil. Firs, birches, alder or aspens dominate in small forests. Broad-leaved forests are common in the southwestern area. It is important to note that the common hornbeam (Carpinus betulus L.) grows naturally in this territory. This is northeastern habitat limit of the species (Navasaitis 2004).

The Lithuanian territory belongs to the Central European forest zone and mixed forest zone. The Lithuanian forests were forming after the last glacial period. Over the last 13 thousand years the forests have been changing from arctic tundra, northern taiga to broad-leaved forests. Pollen spectrum analysis has shown that the Lithuanian forests amounted to 50% of broad-leaved forests, especially oak woodlands, 7500 years ago. During the earlier historic periods the forests covered the major part of today‘s country. It has been estimated that in the 11th - 13th century the forests reached 55%, wetlands and water bodies 23%, and 22% belonged to the agrarian landscape (Kairiūkštis 2003). However, this proportion has changed drastically due to an intensified human economic activity during the last century. Recently the Lithuanian forests cover 19 677 square kilometers and provide 31.7 % of its area. Coniferous woods are 58.8%, the soft leafy forests 35.8%, whereas the hard ones amount to 4.5% of all forests area (Lithuanian Forest Statistics 2004). In southwestern and central Lithuanian Republic the dominating primeval broad-leaved forests have been almost destroyed. The major part has been planted or is "semi-natural". Agricultural lands, covering 53% of all country’s area, have been formed after cutting down the forests and draining the larger half of the marshes (Kurlavičius 2010). The Lithuanian grasslands and natural pastures provide 11% of the agricultural lands or cover more than 47 580 square kilometers. The wetlands cover 5-6% of the Lithuanian territory, whereas rivers, lakes and dams only 4%.

Climate

The Lithuanian climate is often influenced by the Atlantic cyclones which bring much humidity and reduce the differences in temperatures. In the west of the country, especially at the Baltic Sea coast, the maritime climate is evident, but its features decrease in the east (Kaušyla 1981). The multi-year average temperature is 6.2°C, the absolute maximum temperature is 37.5°C and the minimum is -49°C. A vegetation period, when the average day temperature does not exceed 10°C, lasts 145 days averagely. According to the precipitation quantity, Lithuania is in a surplus drainage zone. The average annual precipitation is 661.5mm. The distribution of precipitation is uneven: in the west the quantity is the largest which reaches approximately 900 mm and it does not exceed 500mm of the annual precipitation quantity in the centre and north of the plains in the country. The coldest annual month is January and the average temperature of this month is -5 - 6°C. The warmest month is July when the average temperature reaches 16°C.

Origin and formation of the fauna

Lithuanian beetle faunal structure and composition are closely related to the formative processes of the landscape which took place in this territory. The first, i.e. the oldest Lithuanian territory’s beetles have the characteristic features to exist in tundra and taiga, glacial relicts. They originated during the Subarctic Period, approximately 12-10 thousand years ago. The following betle species could be referred as glacial relicts: Miscodera arctica, Blethisa multipunctata, Pycnoglypta lurida, Oxyporus mannerheimi, Ceruchus chrysomelinus, Ditylus laevis, Trogosoma desparium, Evodinus borealis, and Semanotus undatus amongst others. These species moved to the north of the country when the climate became warmer, while mixed and leafy forests began to form in the Lithuanian territory. During that time eastern Europe and western Siberia species related to pine and small-leaved forests came into existence in the Lithuanian beetle fauna. Approximately 6 thousand years ago the Lithuanian climate became humid, broad-leaved tree species were proliferating as well as the following species belonging to the west European fauna, for instance Calosoma sycopantha, Carabus intricatus, Carabus coriaceus, Dorcus paralelepipedus, Lucanus cervus, Osmoderma barnabita, Liocola marmorata, Gnorimus variabilis, Stenogostus rufus, Elater ferrugineus, Crepidophorus mutilatus, Cucujus cinnaberinus, Cucujus haematodes, Neatus picipes, Platydema violacea, Colydium elongatum, and Cerambyx cerdo. Today there are only small fragments of natural and primeval forests in an agricultural and urban landscape or placesdifficult to approach; therefore, almost all species have become rare and are frequently referred as relicts of the Atlantic Period. The beetle species, ecologically related to open biotypes, can be considered as the youngest in Lithuania. When the area of forests decreased and open habitation expanded the species characteristic to forest steppe and steppe areas proliferated in Lithuania, such as Diachromus germanus, Harpalus hirtipes, Harpalus froelichi, Copris lunaris, Othophagus sp., Hoplia graminicola, Agriotes lineatus, Tytthaspis sedecimpunctata, Phytoecia virgula, Chrysolina aurichalcea, Baris artemisia and similar ones. Beetle species associated with nutritional relations to the increasing number of cultural plants variety found their way into Lithuania while its agriculture was developing. Such species are: Leptinotarsa decemlineata, Phaedon armoraciae, Phyllotreta armoraciae, Psylliodes chlorosephala, Bruchus rufimanus, Tanymecus palliatus, and Ceutorhynchus pleurostigma.

International tourism and trade has been developing, and thanks to it some species have spread in Lithuania: Alphitobius diaperinus, Trogoderma angustum, Rhyzopertha dominica, Attagenus smirnovi, Lyctus africanus, Glischrochilus quadrisignatus, Bruchus pisorum, Acanthoscelides obtectus, and Sitophilus oryzae.

There are no natural barriers preventing the beetle migration in the Lithuanian territory. Therefore, it is evident that the Lithuanian fauna is still in the process of formation. Over the last ten years a number of beetles specific to southern-middle Europe have been discovered, for example Leistus rufomarginatus, Tachyura parvula, Acupalpus luteatus, Elater ferrugineus, Isoriphis melasoides, Melasis buprestoides, Xylotrechus pantherinus, Oulema tristis, and Magdalis caucasica. However, the assumptions must be proved the origin of these species in Lithuania was determined by the climate or whether they are the relicts of the Atlantic Period.

Conservation

The ways to prevent the extinction of plant and animals species, or at least to predict this process and control it, have been sought worldwide for long time. So far, most of the species have disappeared without our knowledge or even without our awareness of their impending departure. The beginning of animal and plant protection can be related to the first nature reserve establishment. In 1937 Žuvintas National Reserve was established on the initiative of a famous scientist, Professor T. Ivanauskas, in order to preserve the unique ecosystem complex of Žuvintas Lake and its surrounding marshes (Kirstukas 2004). Despite that, the list of rare and endangered species was first considered only in 1959 after the first Nature Protection Law had been enacted. In 1996 the Committee for Protection of Nature approved the list of protected plants species compiled by the Lithuanian Academy of Science’s Institute of Botany and Society of Botany. It includes 176 rare or endangered vascular plant species. Following the advice of the Committee for the Protection of Nature, the former Council of Ministers of the Lithuanian SSR established the Lithuanian SSR Red Book in 1976 , standardising the listing procedures. The Lithuanian Red Data Book was published in 1981. It lists 41 species of animals, the complete genus of bumblebees (Bombus), as well as 30 species of vascular plants (Jezerskas et al. 1981). However only one beetle species (Lucanus cervus) was included in the this Red Data Book as a very rare and relict species. Since 1990, Environmental Protection Department of the Republic of Lithuania has been responsible for the compilation and updating of the second edition of the Lithuanian Red Data Book. Overall 102 insect species have been listed in this edition, of which 17 were beetles (Balevičius 1992). Regarding the Lithuanian biodiversity research and its conservation, the Lithuanian situation changed after gaining its independence and signing the Rio de Janeiro Declaration, as well as after the ratification of the International Convention on Biological Diversity (CBD). In order to solve the problems of the protection of the most valuable and endangered species and their habitations, the Birds Directive (1979) was approved, the Network of Protected Areas (Natura 2000) was established, the Habitat Directive (1992), the European Community Strategy on Biological Diversity (1998) and other EU legislative acts and initiatives were implemented in Europe. That fact influenced the separation of valuable territories, regarding their biological diversity, which were given the status of protected areas. According to the High Conservation Values Forests (HCVF) concept, the Lithuanian forests, covering the area of 264.28 square kilometers, were acknowledged to have an exceptional conservation value when the Inventory Project of Woodland Key Habitat was under way in 2000-2005 (Andersson et al. 2005).

The Lithuanian protected territorial system consists of three nature and cultural reserves, one biosphere reserve, five national parks, 30 regional parks and 254 national reserves. The protected territorial system, including nature monuments and nationally protected nature heritage monuments, cover about 7.86 thousand square kilometres (approximately 12% of the country‘s territory) (Kirstukas 2004).

The list of protected insect species has been supplemented substantially in 2003 and a new edition of the Lithuanian Red Data Book has been published (Rašomavičius 2007). Altogether 767 species of animals, plants, fungi and lichens have been included in this book. From the 128 listed insect species, 32 belong to the order Coleoptera (Table 1); however these species have been included in latest Red Data Book on the basis of the status of the species in the list of protected animals of Natura 2000; some of them, such as Ostoma ferruginea Graphoderus bilineatus, Dytiscus latissimus, and Agonum ericeti are not very rare or endangered in Lithuania. At the same time this book does not include such species as Calosoma auropunctatum, Velleius dilatatus, Emus hirtus, Hyleocoetus flabellicornis, Metoecus paradoxus, Tragosoma depsarium, etc., which are rare in the territory of Lithuania and in need of protection.

History of Coleoptera research

Research on Lithuanian Coleoptera does not have a long history. The first records of beetles found in Lithuania were noted by Eduard von Eichwald at the beginning of the 19th century (Eichwald 1830). In his book Zoologia Specialis, he mentioned approximately 150 species of beetles. Some beetle species were mentioned in other works of that century (Wankowicz 1869; Lindeman 1871; Lentz 1879; Osterloff 1889; Seidlitz 1891; Vozhev 1892), but some doubts concerning their authenticity exist since exact locations of the finds were not indicated and the territory of Lithuania in the 19th century and today differ. Some information about Lithuania’s insect fauna can be found in the works published at the beginning of the 20th century. In 1903, Lucas von Heyden released data regarding beetles collected in the environs of Kaunas - 205 species of beetles were reported for Lithuania, 158 of which were recorded for the first time. Ján Roubal (1910) published data on beetles collected in the environs of Vilnius in 1904-1907; 160 species of beetles were included on this list, 120 of which were noted for the first time in Lithuania. Some insect species were also attributed to the Lithuanian fauna in works published in neighbouring countries (Lomnicki 1913; Jakobson 1905-1915). Here, 67 species of insects were noted for Lithuania for the first time. More significant research work concerning the insect fauna of Lithuania was carried out in Vilnius University during the interwar period. However, particular attention was only paid to certain families of beetles such as Haliplidae, Dytiscidae, Carabidae, Rhysodidae, Hydrophilidae, Hydreanidae (Ogijewicz 1933), Silphidae (Kopyłówna 1935), Buprestidae, Elateridae (Ogijewicz 1939), Cerambycidae (Zawadzki 1936; Stanilisówna 1939) and Chrysomelidae (Kamiński 1936). Additionally, quite detailed results of faunistic research carried out in the eastern part of present-day Lithuania were published by Gabriela Mazurowa and Edward Mazur (1939). During this period, besides faunistic research into Coleopteran fauna, research in applied entomology related to agricultural and forest pests were also pursued (Ivanauskas and Vailionis 1922; Mastauskis 1925, 1927, 1936; Ogijewicz 1929, 1931, 1932, 1934, 1938). By 1940, in the current territory of Lithuania, 981 species of beetles had been recorded. During the post-war period from 1945 to 1955, little data concerning the Lithuania fauna was published. Records of 18 species of beetles were mentioned for Lithuania for the first time in the works of Prüffer (1948), (Horion (1949, 1955), (Richter (1949, 1952), and Medvedev (1952). Focused beetle research has been conducted by Stanislovas Mastauskis and Simonas Pileckis since 1955 in the Department of Plant Protection, Lithuanian Academy of Agriculture (currently the University). Lithuania’s beetle species list was considerably enlarged after the beetle collections of entomologist Alfonsas Palionis were analysed. Based on Palionis’ data, 351 beetle species new for Lithuania were registered. Moreover, important works concerning harmful beetles in Lithuanian forests were published; these including nine species of bark beetles (Curculionidae: Scolytinae) (Mastauskis and Pileckis 1959) and four species of jewel beetles (Buprestidae) (Pileckis 1958a) recorded for the first time in Lithuania. Pileckis (1960), who composed the first list of Lithuanian beetles, made a great contribution to the exploration of the fauna. The author presented a critical analysis of previous publications regarding Lithuanian beetles and summarised results of the audit of beetle collections held in natural history museums of Lithuania and in private collections. In the first list of Lithuanian beetles, 1446 beetle species were listed, 122 of which were for the first time. It should be noted that Pileckis (1960) not only notified new beetle species for Lithuania, but also proposed to remove some species from the list of misidentified beetles such as Lethrus apterus, Aphodius alpinus, Anoplodera sequensi, Clytus rhamni, Isotomus speciosus and Acanthocinus reticulatus. In the years after 1960, numerous works of faunistic and applied entomology were published (Pileckis 1962, 1963a, b, 1968a, b, 1970a, b, 1972; Lešinskas and Pileckis 1967; Pileckis et al. 1968; Zajančkauskas and Pileckis 1968; Kazlauskas and Pileckis 1968; Mastauskis 1970; Valenta and Jakaitis 1972; Jakaitis 1973; Sharova and Grüntal 1973; Mensonienė 1974) and these were summarised in the first monograph on Lithuanian beetles (Pileckis 1976a). In the monograph, the body composition of beetles, reproductive and developmental characteristics, environmental and distribution peculiarities, regularities of fauna formation, description of research process, as well as identification keys for some beetle groups and a systematic list of Lithuanian beetles are presented. 2203 beetle species were included on this list, 436 of which were mentioned for for the first time. It is worth noting that the 1970s were the most productive period in terms of beetle fauna research in Lithuania: 569 beetle species were noted the first time (Figure 1). It is also worth noting that besides the above-mentioned works, a Prussian beetle list was also published in the same decade. On this list, beetles found in Klaipėda region and near Juodkrantė in the Curonian Spit were noted (Bercio and Folwaczny 1979); 84 beetle species were new for Lithuania. In the 1980s, Vidmantas Monsevičius played a very important role in faunistic research, aiming to determine the species composition of Staphylinidae. During this period, 210 species of rove beetles new for Lithuania were recorded (Monsevičius 1982, 1883, 1985, 1986a, b, 1987, 1988a, b; Monsevičius and Jakaitis 1984). Beetles living in forest ecosystems were also actively studied; 108 new beetle species were found (Perkovskij and Monsevičius 1982; Pileckis and Monsevičius 1982; Pileckis and Jakaitis 1982, 1989; Slavinskas 1982; Pileckis 1984, 1988; Jakaitis 1985; Gaidienė and Ferenca 1988; Ferenca 1988). A significant contribution to the knowledge of the Lithuanian fauna was also made by the Estonian entomologist Georg Miländer; in his work, 42 beetle species new for Lithuania were mentioned (Miländer 1982; Miländer et al. 1984).

A more comprehensive review of faunistic records of beetles in Lithuania was provided by authors of subsequent volumes of the Lietuvos Fauna. Vabalai [Lithuanian Fauna. The beetles] (Pileckis and Monsevičius 1995 and 1997). This monograph consists of two parts, general and systematic. The first deals with the body structure and bionomics of beetles, the history of coleopteran research in Lithuania, stages of beetle fauna formation during the Postglacial Period, peculiarities of coleopteran distribution in the Republic as well as their significance. In the second part the descriptions of families of Coleoptera, genera and some species as well as distribution (in five regions) and occurrence rate (within 5 degrees) of each species are given. The monograph includes information on 2895 beetle species; 133 species are noted the first time for Lithuania. It is important to note that this is the first attempt to forecast expected beetle species in Lithuania: a number of species expected for each genus is given, totalling more than 990 additional species. Some valuable faunistic information about can be found in the Catalogue of Entomological Collections of Tadas Ivanauskas Zoological Museum. Therein 75 species of beetles have been recorded for the first time in Lithuania. Some information on new beetle species found in Lithuania can also be found in the catalogues of neighbouring countries (Silfverberg 1992, 2004; Lundberg and Gustafsson 1995; Althoff and Danilevsky 1997). Moreover, some species of beetles have been noted only in these catalogues and are not confirmed by more circumstantial information. The same can be said of later catalogues of faunistic works such as the Catalogue of Palaearctic Coleoptera and Fauna Europaea Web Service. In total, 122 species of beetles are noted the first time.

The publication of the most recent monograph (Pileckis and Monsevičius 1995 and 1997) induced an increase in faunistic investigations in Lithuania, which resulted in more than 50 publications containing new faunistic data on beetles the most important are arranged in the table 2.

Methods and conventions

In this check-list we use the classification of the order Coleoptera accepted by Bouchard et al. (2011), the genera and species names we use following accepted ones in the Catalogue of Palaearctic Coleoptera (Löbl and Smetana (eds.) 2003-2010) with some details accepted by other authors: Staphylinidae by Herman (2001), Buprestoidea by Kuban et al. (2006), Tenebrionoidea by Silfverberg (2004), Cerambycidae by Althoff and Danilevsky (1997), and Curculionoidea by Alonso-Zarazaga and Lyal (1999). The main taxonomic categories referred to in this check-list are the following: order, suborder, family, subfamily, tribe, genus and species. Higher taxonomic categories, like series, super- or sub-tribes, are omitted (for details see Bouchard et al. (2011)). Most common synonyms and subspecies names are included, especially the names used in the two volumes Lithuanian monograph (Pileckis and Monsevičius 1995, 1997). The list of Lithuanian beetles was compiled on the basis of published literature sources and revisions of public beetle collections. The catalogues, checklists and other faunistic literature of neighbouring countries have been used to compile the list of beetles species expected in Lithuanian fauna. The species are arranged alphabetically in their respective genera. Expected for Lithuania species names are placed in square brackets […]. The species excluded from the list of Lithuanian beetles are marked by asterisk (*). The officially protected species, included in the Red Data Book of Lithuania (2007) are marked by the abbreviation RDB. The comments (#) are presented after the list and referenced by sequential numbers.

Results and discussion

All available published and interactive sources on beetles found in Lithuania have been summarized in the current study. In addition to verified records, 166 species noted for Lithuania only in several European catalogues (Enumeratio Coleopterorum Fennscandiae, Daniae et Baltiae (Silfverberg 1992, 2004), Catalogue of Palaearctic Coleoptera (Löbl and Smetana (eds.), 2003-2010), Fauna Europaea Web Service (2001-2011)) and other works of foreign scientists (Osterloff 1889; Jakobson 1905–1915; Łomnicki 1913; Lundberg and Gustafsson 1995; Althoff and Danilevsky 1997) are included in this new list of Lithuanian beetles (Table 3). The primary sources of these findings remains unknown to the authors. Another 17 species were noted for Lithuania by native authors, but this data is untrustworthy as the collected material is lost and confirmation remain absent.

For reasons of clarity, 19 previously noted but later excluded species are included in the current checklist with comments. Moreover, we recommend the removal of 43 species from the list as they were misidentified and erroneously reported in previous works or are species which have been noted by mistake and their distribution in Lithuania is untrustworthy (Table 4).

As a result, 3597 species of beetles are included in the new checklist. Based on the faunal data from neighbouring countries (Lundberg and Gustafsson 1995; Silfverberg 2004; Telnov 2004, 2010; Barševskis 2003; Barševskis et al. 2005; Alexandrovitch et al. 1996; Szujecki 1961, 1976; Stebnicka 1976, 1978; Smrezyński 1965, 1966, 1968, 1972, 1974, 1976; Burakowski et al. 1973, 1974, 1976, 1978, 1979, 1980, 1981, 1986a, b, c, 1987, 1989, 1990, 1991; Wanat and Mokrzycki 2005, Tarnawski and Buchholz 2008a, b; Süda 2006, 2009; Fauna Europaea Web Service (2009) and other faunistic works), a further 1390 species of beetles are included as expected species for Lithuanian fauna (Table 5).

The analysis of faunal data in Lithuania and neighbouring countries provides evidence that the Lithuanian coleopteran fauna has yet to be completely investigated and it is estimated that approximately 28% of beetle species remain undiscovered in Lithuania. More than 85% of expected beetle species have been found in the following families: Cerylonidae, Geotrupidae, Haliplidae, Kateridae, Lycidae, Lucanidae, Mycetophagidae, Scarabaeidae and Silphidae. In families with few species such as Alexiidae, Boridae, Byturidae, Dascilidae, Drilidae, Eucinetidae, Lampyridae, Lymexilidae, Megalopodidae, Nemonychidae, Nosodendridae, Noteridae, Orsodacnidae, Pyrochroidae, Pythidae, Psephenidae, Rhysodidae, Sphaeritidae, Sphaeriusidae, Sphindidae, Stenotrahelidae, and Trogidae all possible species have been already discovered. However in some beetle families such as Aderidae, Bothrideridae, Eucnemidae, Laemoploeidae, Mordellidae, Ptiliidae, Scraptidae and Throscidae less than 50% of all possible species are known. At present, the beetle species belonging to 92 familiesare reported for Lithuania and the species from the other 9 families such as Agyrtidae, Biphylidae, Deradontidae, Mycteridae, Ochodaeidae, Phleophilidae, Phloeostichidae, Prostomidae, Trachypachidae are expected.

List of species

MYXOPHAGA Crowson, 1955.

SPHAERIOIDEA Erichson, 1845.

SPHAERIUSIDAE Erichson, 1845 = MICROSPORIDAE Crotch, 1873. (Sphaeriidae)

Sphaerius Waltl, 1838 = Microsporus Kolenati, 1846.

acaroides Waltl, 1838 = obsidianus (Kolenati, 1846). Pileckis and Monsevičius 1995; Silfverberg 2004; Löbl 2009.

ADEPHAGA Crowson, 1955.

GYRINIDAE Latreille, 1810.

Gyrininae Latreille, 1810.

Gyrinini Latreille, 1810.

Aulongyrus Régimbart, 1883.

[concinnus (Klug, 1834)]. Known in northeastern Poland (Burakowski et al. 1976).

Gyrinus O.F. Müller, 1764.

aeratus Stephens, 1835. Pileckis and Monsevičius 1995; Monsevičius 1998; Barševskis 2001a; Silfverberg 2004; Barševskis et al. 2005; Alekseev 2010a.

caspius Ménétriés, 1832. Barševskis 2001a; Mazzoldi 2003, 2009; Silfverberg 2004; Alekseev 2010a.

[colymbus Erichson, 1837]. Known in Kaliningrad region (Alekseev 2010a), eastern Belarus (Alexandrovitch et al. 1996) and Denmark (Mazzoldi 2009).

distinctus Aubé, 1836. Ogijewicz 1933; Pileckis 1960, 1976a, b, 1979; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Mazzoldi 2003; Alekseev 2010a.

marinus Gyllenhal, 1808. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Kovács et al. 2008; Alekseev 2010a.

minutus Fabricius, 1798. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Alekseev 2010a.

natator (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Gliaudys 2001; Ferenca 2006b; Mazzoldi 2009; Alekseev 2010a.

opacus C.R. Sahlberg, 1819. Barševskis 2001a; Silfverberg 2004; Alekseev 2010a.

paykulli G. Ochs, 1927. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Šablevičius 2003a; Alekseev 2010a.

pullatus Zaitzev, 1908. Barševskis, 2001; Silfverberg 2004; Alekseev 2010a.

substriatus Stephens, 1828. Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

[suffriani W. Scriba, 1855]. Known in Baltic Sea coast region in Poland (Burakowski et al. 1976), Kaliningrad region (Alekseev 2010a), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Mazzoldi 2009), Estonia (Rutanen 2004).

urinator Illiger, 1807. Pileckis 1960, 1976a, b, 1979; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Barševskis et al. 2005; Ferenca 2006b; Alekseev 2010a.

Orectochilini Régimbart, 1882.

Orectochilus Dejean, 1833.

villosus (O.F. Müller, 1776). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Mazzoldi 2003, 2009; Kovács et al. 2008; Alekseev 2010a; Šablevičius 2011.

TRACHYPACHIDAE Thomson, 1857.

Trachypachinae Thomson, 1857.

Trachypachus Motschulsky, 1844.

[zetterstedtii (Gyllenhal, 1827)].Known in Latvia (Barševskis 2003).

RHYSODIDAE Laporte, 1840.

Rhysodini Laporte, 1840.

Rhysodes Germar, 1822.

sulcatus (Fabricius, 1787).# 1. Ogijewicz 1933; Pileckis 1960.

CARABIDAE Latreille, 1802.

Nebriinae Latreille, 1802.

Nebriini Laporte, 1834.

Leistus Frölich, 1799.

ferrugineus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997, 2001; Šablevičius 2000b; Gliaudys 2001; Farkač and Janata 2003; Tamutis 2005a; Ferenca 2006b; Tamutis et al. 2007; Alekseev 2008a, c; Vigna Taglianti 2009.

piceus Frölich, 1799. Šablevičius and Ferenca 1995; Silfverberg 1996, 2004; Monsevičius 1998; Šablevičius 2000a, b, 2003a, b; Farkač and Janata 2003; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

rufomarginatus (Duftschmid, 1812). Barševskis 2001a, 2003; Ferenca et al. 2002; Silfverberg 2004; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009; Ivinskiset al. 2009.

terminatus (Panzer, 1793) = rufescens (Fabricius, 1775) nec (Ström, 1768).Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997, 2001; Farkač and Janata 2003; Žiogas and Zolubas 2005; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Nebria Latreille, 1802.

brevicollis (Fabricius, 1792).Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca et al. 2002; Farkač and Janata 2003; Žiogas and Zolubas 2005; Ferenca 2006b; Žiogas and Vaičikauskas 2007a, b; Alekseev 2008c; Vigna Taglianti 2009.

livida (Linnaeus, 1758). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Ferenca 2006b; Alekseev 2008c.

rufescens (Ström, 1768) = gyllenhali (Schönherr, 1806). # 2. Pileckis 1968b, 1970a, 1976a, b, 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2008c.

[salina Fairmaire & Laboulbène, 1854]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Silfverberg 1992, 2004).

Notiophilini Motschulsky, 1850.

Notiophilus Duméril, 1806.

aesthuans Dejean, 1826 = pusillus Waterhouse, 1833 nec (Schreber, 1759).Ogijewicz 1933; Pileckis 1960, 1970a, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Barševskis 2001; Bousquet and Barševskis 2003; Alekseev 2005; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

aquaticus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Bousquet and Barševskis 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

biguttatus (Fabricius, 1779). Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Bousquet and Barševskis 2003; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

germinyi Fauvel, 1863 = stipraisi Barševskis, 1993.Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Barševskis 2001a; Bousquet and Barševskis 2003; Alekseev 2008c; Vigna Taglianti 2009.

palustris (Duftschmid, 1812). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis and Zolubas 2001; Bousquet and Barševskis 2003; Tamutis et al. 2004, 2007; Tamutis 2005a; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Dapkus and Tamutis 2007; Alekseev 2008c; Vigna Taglianti 2009.

[rufipes Curtis, 1829]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark, Poland, Estonia, Kaliningrad region (Vigna Taglianti 2009).

Pelophilini Kavanaugh, 1996.

Pelophila Dejean, 1821.

[borealis (Paykull, 1790)]. Known in Estonia (Haberman 1968); Latvia (Barševskis 2003), central Sweden (Lundberg and Gustafsson 1995), northern Belarus (Alexandrovitch et al. 1996).

Cicindelinae Latreille, 1802.

Cicindelini Latreille, 1802.

Cicindela Linnaeus, 1758.

campestris Linnaeus, 1758. Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Gliaudys 2001; Puchkov and Matalin 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

hybrida Linnaeus, 1758. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Gliaudys 2001; Puchkov and Matalin 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

RDB maritima Latreille & Dejean, 1822. Pileckis 1963b, 1976a, b, 1979; Bercio and Folwaczny 1979; Balevičius 1992; Šablevičius 1995, 2011; Pileckis and Monsevičius 1995; Ferenca 2004; 2006a, b; Ivinskis and Rimšaitė 2005; Tamutis 2005b; Rašomavičius 2007; Alekseev 2008c, 2010b; Ivinskis et al. 2009; Vigna Taglianti 2009.

sylvatica Linnaeus, 1758. Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Alekseev 2003, 2008c; Puchkov and Matalin 2003; Ferenca 2006b; Vigna Taglianti 2009; Šablevičius 2011.

Cylindera Westwood 1831.

arenaria viennensis (Schrank, 1781). Ferenca et al. 2002; Barševskis 2003; Ivinskis et al. 2004a; Alekseev 2008c; Vigna Taglianti 2009.

germanica (Linnaeus, 1758).Eichwald 1830 (Cicindela); Pileckis 1960, 1976a (Cicindela); Silfverberg 1992, 2004; Gaidienė 1993 (Cicindela); Pileckis and Monsevičius 1995 (Cicindela); Gliaudys 2001 (Cicindela); Inokaitis 2004; Tamutis 2005b, Tamutis et al. 2007; Alekseev 2008c; Vaivilavičius 2008; Vigna Taglianti 2009.

Carabinae Latreille, 1802.

Carabini Latreille, 1802.

Calosoma Weber, 1801.

auropunctatum (Herbst, 1784). Pileckis 1968a, 1976a, 1982; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 1999, 2000, 2005a; Gliaudys 2001; Bousquet et al. 2003; Tamutis et al. 2007; Alekseev 2008c; Vigna Taglianti 2009.

RDB inquisitor (Linnaeus, 1758). Pileckis 1968a, 1976a, b, 1979, 1982; Bercio and Folwaczny 1979; Balevičius 1992; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Pileckis and Vengeliauskaitė 1996; Šablevičius 2000a, 2011; Gliaudys 2001; Ehnström et al. 2003; Bousquet et al. 2003; Ferenca 2004, 2006a; Ivinskis et al. 2004b, 2006, 2007; Tamutis 2005b; Rašomavičius 2007; Kriaučiūnienė and Zaplatkin 2007; Alekseev 2008c, 2010b; Vigna Taglianti 2009; Bačianskas 2009; Noreika 2009.

[investigator (Illiger, 1798)]. Known in southern Sweden (Lundberg and Gustafsson 1995), northeastern Poland (Burakowski et al. 1973), Kaliningrad region (Bercio and Folwaczny 1979, Vigna Taglianti 2009).

RDB sycophanta (Linnaeus, 1758).# 3. Lentz 1879; Pileckis 1976a; Bercio and Folwaczny 1979; Pileckis and Monsevičius 1982, 1995; Balevičius 1992; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Vengeliauskaitė 1996; Gliaudys 2001; Ehnström et al. 2003; Bousquet et al. 2003; Tamutis 2005b; Rašomavičius 2007; Alekseev 2008c, 2010b; Vigna Taglianti 2009; Šablevičius 2011.

Callisthenes Fischer von Waldheim 1821.

[reticulatum (Fabricius, 1787)]. # 4. Known in southern Sweden (Lundberg and Gustafsson 1995), northeastern Poland (Burakowski et al. 1973), western Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Bercio and Folwaczny 1979).

Carabus Linnaeus, 1758.

arcensis Herbst, 1784 = arvensis Herbst, 1784.Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Gedminas and Žiogas 2001, 2007; Bousquet et al. 2003; Ferenca 2006b; Lynikienė 2006; Žiogas et al. 2006; Žiogas and Vaičikauskas 2007b; Dapkus and Tamutis 2007, 2008a; Alekseev 2008c; Vigna Taglianti 2009.

[auratus Linnaeus, 1761]. Known in northeastern Poland (Burakowski et al. 1973), Denmark (Vigna Taglianti 2009).

cancellatus tuberculatus Dejean, 1826. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Zajančkauskas and Pileckis 1968; Strazdienė 1976; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Gliaudys 2001; Bousquet et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Lynikienė 2006; Žiogas and Vaičikauskas 2007b; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009; Šablevičius 2011.

clathratus Linnaeus, 1761. Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Bousquet et al. 2003; Ferenca 2003, 2006b; Ivinskis et al. 2004a; Alekseev 2008c; Vigna Taglianti 2009.

convexus Fabricius, 1775. Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a, 1982; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ferenca et al. 2002; Šablevičius 2003a, b; Bousquet et al. 2003; Žiogas and Zolubas 2005; Alekseev 2008c; Ivinskis et al. 2009; Vigna Taglianti 2009; Noreika 2009.

RDB coriaceus Linnaeus, 1758. Eichwald 1830; Jakobson 1905-1915; Pileckis 1960, 1976a, b, 1979, 1982; Lešinskas and Pileckis 1967; Strazdienė 1981, 1988; Balevičius 1992; Gaidienė and Ferenca 1992; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ivinskis et al. 1996b, 1997b, 1998b, 2000, 2004a, 2006, 2009; Ferenca 1996, 2006b; Auglys 1998; Gliaudys 2001; Bousquet et al. 2003; Tamutis 2005b; Rašomavičius 2007; Vaivilavičius 2008; Sruoga 2008; Alekseev 2008c, 2010b; Dapkus & Tamutis 2009; Vigna Taglianti 2009; Šablevičius 2011.

glabratus Paykull, 1790.Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Bousquet et al. 2003; Ferenca 2006b; Lynikienė 2006; Žiogas and Vaičikauskas 2007b; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

granulatus Linnaeus, 1758. Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a, 1982; Zajančkauskas and Pileckis 1968; Strazdienė 1981; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Gliaudys 2001; Bousquet et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

hortensis Linnaeus, 1758. Eichwald 1830; Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Bousquet et al. 2003; Žiogas and Zolubas 2005; Lynikienė 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007a, b; Dapkus and Tamutis 2007, 2008a; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

RDB intricatus Linnaeus, 1761. Ferenca 1996; Ivinskis et al. 1996a, 1997b, 1998b, 2000, 2004a; Jonaitis et al. 2000; Šablevičius 2000b; Ferenca et al. 2002; Tamutis 2005b; Rašomavičius 2007; Alekseev 2008c, 2010b.

[marginalis Fabricius, 1794]. Known in northeastern Poland (Burakowski et al. 1973), eastern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

menetriesi (Faldermann in Hummel, 1827). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Balevičius 1992; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet et al. 2003; Tamutis 2005b; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008, 2009; Alekseev 2008c; Vigna Taglianti 2009.

nemoralis O.F. Müller, 1764. Ogijewicz 1933; Pileckis 1960, 1976a, 1982; Zajančkauskas and Pileckis 1968; Strazdienė 1981; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Bousquet et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

RDB nitens Linnaeus, 1758. Ogijewicz 1933; Pileckis 1960, 1976a, 1982; Zajančkauskas and Pileckis 1968; Balevičius 1992; Silfverberg 1992, 2004; Gaidienė 1993; Obelevičius 1994; Ostrauskas and Kubertavičius 1994; Šablevičius 1994, 2000b, 2003a, b, 2011; Pileckis and Monsevičius 1995; Ivinskis et al. 1996b, 2000, 2004a; Pankevičius 2000, 2007; Monsevičius 1997, 2001; Gliaudys 2001; Bousquet et al. 2003; Meržijevskis 2004; Dapkus 2004; Tamutis 2005b; Ferenca 2006b; Lopeta 2007a; (misprint as Callisthenes coriaceus L. in Lopeta 2007b); Rašomavičius 2007; Uselis et al. 2007; Butvila et al. 2007; Alekseev 2008c, 2010b; Vigna Taglianti 2009.

[problematicus harcyniae Sturm, 1815].Known in northeastern Poland (Burakowski et al. 1973), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009).

*scheidleri Panzer, 1799. # 5. Pileckis 1960.

violaceus andrzejuscii Fischer von Waldheim, 1823. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a, 1982; Strazdienė 1988; Silfverberg 1992, 2004; Ferenca et al. 2002; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Bousquet et al. 2003; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Dapkus and Tamutis 2007; Vaivilavičius 2008; Sruoga 2008; Alekseev 2008c; Vigna Taglianti 2009; Šablevičius 2011.

Cychrini Laporte, 1834.

Cychrus Fabricius, 1794.

caraboides (Linnaeus, 1758) = rostratus (Linnaeus, 1761). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Monsevičius 2001; Gliaudys 2001; Häckel 2003; Žiogas and Zolubas 2005; Lynikienė 2006; Ferenca 2006b; Dapkus and Tamutis 2007; Žiogas and Vaičikauskas 2007a, b; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Loricerinae Bonelli, 1810.

Loricerini Bonelli, 1810.

Loricera Latreille, 1802.

pilicornis (Fabricius, 1775) = caerulescens auct. nec (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Strazdienė 1976; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Bousquet 2003a; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Žiogas et al. 2006, 2009; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Omophroninae Bonelli, 1810.

Omophronini Bonelli, 1810.

Omophron Latreille, 1802.

limbatum (Fabricius, 1776). Eichwald 1830; Pileckis 1960, 1976a; Gaidienė and Ferenca 1992; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hůrka 2003; Ferenca 2004, 2006b; Vaivilavičius 2008; Alekseev 2008a, c; Vigna Taglianti 2009; Ivinskis et al. 2009, Valainis 2010; Šablevičius 2011.

Elaphrinae Latreille, 1802.

Elaphrini Latreille, 1802.

Blethisa Bonelli, 1810.

multipunctata (Linnaeus, 1758). Eichwald 1830; Jakobson 1905-1915; Pileckis 1960, 1970a, 1976a, b, 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Goulet 2003; Šablevičius 2003a; Ivinskis et al. 2004a; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

Elaphrus Fabricius, 1775.

[angusticollis R.F. Sahlberg, 1844]. Known in northwestern Belarus (Alexandrovitch et al. 1996), Latvia (Barševskis 2003).

aureus O.F. Müller, 1821. Pileckis 1968b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Gedminas et al. 1996; Ferenca et al. 2002; Goulet 2003; Alekseev 2008c; Vaivilavičius 2008; Vigna Taglianti 2009.

cupreus Duftschmid, 1812. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis and Zolubas 2001; Goulet 2003; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

riparius (Linnaeus, 1758). Eichwald 1830; Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1968b, 1970a, 1976a, b, 1979; Lešinskas and Pileckis 1967; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Goulet 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

uliginosus Fabricius, 1792. Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Goulet 2003; Alekseev 2008c; Vigna Taglianti 2009.

Scaritinae Bonelli, 1810.

Clivinini Rafinesque 1815.

Clivina Latreille, 1802.

collaris (Herbst, 1784). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Balkenohl 2003; Ferenca 2006b; Alekseev 2008c.

fossor (Linnaeus, 1758).Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Strazdienė 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Balkenohl 2003; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

Dyschirius Bonelli, 1810.

angustatus (Ahrens, 1830). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ferenca 2006b; Alekseev 2008c.

digitatus (Dejean, 1825). Pileckis 1963b, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2008c.

obscurus (Gyllenhal, 1827). Pileckis 1962, 1963b, 1970a, 1976a, b, 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Alekseev 2008a, c.

thoracicus (P. Rossi, 1790) = arenosus Stephens, 1827. Eichwald 1830; Jakobson 1905-1915; Roubal 1910; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Alekseev 2008a, c.

Dyschiriodes Jeannel 1941.

aeneus (Dejean, 1825).Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Dyschirius); Monsevičius 1997; Alekseev 2008c.

[chalceus (Erichson, 1837)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1973), Kaliningrad region, Estonia, Denmark (Vigna Taglianti 2009).

globosus (Herbst, 1784). Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Dyschirius); Monsevičius 1997; Tamutis 2000; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c.

impunctipennis (Dawson, 1854). Pileckis 1976a, b, 1979; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dyschirius); Alekseev 2008c.

intermedius (Putzeys, 1846). Lentz 1879; Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dyschirius); Tamutis 2000, 2005a; Tamutis et al. 2004; Alekseev 2008c; Ivinskis et al. 2009.

laeviusculus (Putzeys, 1846). Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dyschirius); Alekseev 2008c.

neresheimeri (Wagner, 1915).Lentz 1879; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dyschirius); Alekseev 2008c.

[nigricornis (Motschulsky, 1844) = helleni (P.W.J.Müller, 1922) = septentrionum (Munster, 1923)]. Known in northern Sweden (Lundberg and Gustafsson 1995), Latvia (Barševskis 2003), Kaliningrad Region, Estonia, (Vigna Taglianti 2009).

nitidus (Dejean, 1825).Eichwald 1830; Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dyschirius); Monsevičius 1997; Ivinskis et al. 1998; Ferenca et al. 2002; Alekseev 2008c.

politus (Dejean, 1825). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Dyschirius); Tamutis 2000, 2005a; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c.

[salinus (Schaum, 1843)]. Known in southern Sweden (Lundberg and Gustafsson 1995), coast of Baltic Sea in Poland (Burakowski et al. 1973), Latvia (Barševskis 2003), Kaliningrad Region, Estonia, Denmark (Vigna Taglianti 2009).

tristis (Stephens, 1827) = luedersi Wagner, 1915. Bercio and Folwaczny 1979; Pileckis and Monsevičius 1982, 1995 (Dyschirius); Silfverberg 1992, 2004; Alekseev 2008c.

Broscinae Hope, 1838.

Broscini Hope, 1838.

Broscus Panzer, 1813.

cephalotes (Linnaeus, 1758). Heyden 1903; Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Ferenca 2006b; Alekseev 2008c.

Miscodera Eschscholtz, 1830.

arctica (Paykull, 1798). Pileckis 1968a, 1970a, 1976a, b, 1979, 1982; Ivinskis et al. 1984, 2004a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca et al. 2002; Alekseev 2008c.

Trechinae Bonelli, 1810.

Trechini Bonelli, 1810.

Perileptus Schaum, 1860.

[aureolatus (Creutzer, 1799)]. Known in Sweden (Lundberg and Gustafsson 1995), northeastern Poland (Burakowski et al. 1973), Latvia (Barševskis 2003), Belarus (Alexandrovitch et al. 1996).

Trechus Clairville, 1806.

austriacus Dejean, 1831. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Moravec et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

obtusus Erichson, 1837.Ferenca et al. 2006, 2007; Ivinskis et al. 2009.

quadristriatus (Schranck, 1781). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Moravec et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Žiogas et al. 2006; Alekseev 2008c; Vigna Taglianti 2009; Ostrauskas and Ferenca 2010.

rivularis (Gyllenhal, 1810). Pileckis and Monsevičius 1982, 1995 (Epaphius); Ivinskis et al. 1984; Silfverberg 1992, 2004; Monsevičius 1997; Moravec et al. 2003; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

rubens (Fabricius, 1792). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 1999, 2000, 2005a; Moravec et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

secalis (Paykull, 1790). # 19. Roubal 1910; Ogijewicz 1933 (Epaphius); Pileckis 1960, 1976a (Epaphius); Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Epaphius); Monsevičius 1997; Tamutis 2000, 2005a; Moravec et al. 2003; Tamutis et al. 2004; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Trechoblemus Ganglbauer, 1891.

micros (Herbst, 1784). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 2000, 2003, 2005a; Moravec et al. 2003; Alekseev 2008c; Vigna Taglianti 2009; Ivinskis et al. 2009.

Blemus Dejean, 1821 = Lasiotrechus Ganglbauer, 1892.

discus (Fabricius, 1792). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000; Moravec et al. 2003; Tamutis et al. 2004; Alekseev 2008c; Vigna Taglianti 2009.

Pagonini Laporte, 1834.

Pogonus Dejean, 1821.

chalceus (Marsham, 1802). # 6. Pileckis 1960, 1976a, b, 1979; Pileckis and Monsevičius 1995; Bousquet 2003b; Ivinskis and Rimšaitė 2005; Ferenca 2006b; Vigna Taglianti 2009.

[luridipennis (Germar, 1822)]. Known in Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1973), Latvia, Denmark, Kaliningrad region (Vigna Taglianti 2009).

Bembidiini Stephens, 1827.

Asaphidion Gozis, 1886.

[austriacum Schweiger, 1975]. Known in Latvia (Barševskis 2003), Belarus (Vigna Taglianti 2009).

caraboides (Schrank, 1781). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Barševskis 2003; Ferenca 2006b; Alekseev 2008c.

[*curtum (Heyden, 1870)].# 7. Tamutis and Ferenca 2006.

flavipes (Linnaeus, 1761). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Marggi et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Alekseev 2008c.

pallipes (Duftschmid, 1812). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Marggi et al. 2003; Tamutis 2005a; Tamutis et al. 2007; Alekseev 2008c.

Odontium LeConte, 1848.

argenteolum (Ahrens, 1812). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

*foraminosum (Sturm, 1825). Marggi et al. 2003; Vigna Taglianti 2009.# 8.

litorale (Olivier, 1790). Heyden 1903; Jakobson 1905-1915; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009; Šablevičius 2011.

striatum (Fabricius, 1792). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

velox (Linnaeus, 1761). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2004, 2006b; Alekseev 2008a, c; Vigna Taglianti 2009.

Metallina Motschulsky, 1850.

lampros (Herbst, 1784). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Tamutis 2005a; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

nigricorne (Gyllenhal, 1827). Jakobson 1905-1915; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Alekseev 2008c.

pygmaea (Fabricius, 1792). Jakobson 1905-1915; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

properans (Stephens, 1828). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Tamutis 1999, 2000, 2005a; Marggi et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Vigna Taglianti 2009; Alekseev 2008c.

Phyla Motschulsky, 1844.

bipunctatum (Linnaeus, 1761). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

obtusa (Audinet-Serville, 1821). Gaidienė 1993; Tamutis 2003; Marggi et al. 2003; Silfverberg 2004; Alekseev 2008c; Ivinskis et al. 2009; Vigna Taglianti 2009.

pallidipenne (Illiger, 1802). Łomnicki 1913; Pileckis 1968b, 1976a, b, 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2004; Ivinskis and Rimšaitė 2005; Alekseev 2008c; Vigna Taglianti 2009.

Princidium Motschulsky, 1864.

punctulatum (Drapiez, 1821). Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1970a, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

ruficolle (Panzer, 1797). Heyden 1903; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

Ocydromus Clairville, 1806.

bruxellensis (Westmaël, 1835) = rupestre auct. nec (Linnaeus, 1758). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

*coeruleus (Audient-Serville, 1821). Ferenca et al. 2002; disproved by Ferenca (2003).

cruciatus polonicum (P.W.J. Müller, 1930). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Šablevičius 2000b; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[dauricus (Motschulsky, 1844)]. Known in Latvia and Estonia (Vigna Taglianti 2009).

*decorus (Panzer, 1799). # 15. Marggi et al. 2003; Vigna Taglianti 2009.

deletus (Audinet-Serville, 1821) = nitidulum (Marsham, 1802) nec (Schrank, 1781). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

femoratus (Sturm, 1825). Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

fluviatilis (Dejean, 1831). # 15. Marggi et al. 2003; Vigna Taglianti 2009.

lunatus (Duftschmid, 1812). Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

[maritimus (Stephens, 1835)]. Known in Denmark and Finland (Vigna Taglianti 2009).

*modestus (Fabricius, 1801). # 15. Marggi et al. 2003; Vigna Taglianti 2009.

monticola (Sturm, 1825). Miländer et al. 1984; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

[obscurellus (Motschulsky, 1844)]. Known in Latvia (Telnov et al. 2006), Estonia, Denmark (Vigna Taglianti 2009), southern Sweden (Lundberg and Gustafsson 1995).

saxatilis (Gyllenhal, 1827).Heyden 1903; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

stephensii (Crotch, 1869). Ferenca 2003; Marggi et al. 2003 Vigna Taglianti 2009; Alekseev 2008c.

tetracolus (Say, 1823) = ustulatus (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Marggi et al. 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Alekseev 2008c.

tetragrammus illigeri (Netolitzky, 1914) = quadriguttatus (Illiger, 1798) = genei auct. nec (Küster, 1847). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

*tibialis (Duftschmid, 1812). # 15. Marggi et al. 2003; Vigna Taglianti 2009.

Sinechostictus Motschulsky, 1864.

*elongatus (Dejean, 1831). # 9. Pileckis 1960 (Bembidion); Ferenca 2006b.

Plataphus Motschulsky, 1864.

[crenulatus (R.F. Sahlberg, 1844)]. Known in Estonia, Latvia (Vigna Taglianti 2009).

[difficilis (Motschulsky, 1844)]. Known in Estonia, Latvia (Vigna Taglianti 2009), central Sweden (Lundberg and Gustafsson 1995).

[hastii (C.R. Sahlberg, 1827)]. Known in Estonia, Latvia (Vigna Taglianti 2009), northern Sweden (Lundberg and Gustafsson 1995).

[prasinus (Duftschmid, 1812)]. Known in Estonia, Latvia (Vigna Taglianti 2009), central Sweden (Lundberg and Gustafsson 1995).

[virens (Gyllenhal, 1827)]. Known in Estonia, Latvia (Vigna Taglianti 2009), central Sweden (Lundberg and Gustafsson 1995).

Notaphus Dejean, 1821.

dentellum (Thunberg, 1787). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

varium (Olivier, 1795).Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Šablevičius 2000b; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

semipunctatum (Donovan, 1806). Roubal 1910; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

obliquum (Sturm, 1825). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Šablevičius 2000b; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[ephippium (Marsham, 1802)].Known in Denmark and Germany (Vigna Taglianti 2009).

Emphanes Motschulsky, 1850.

azurescens (Dalla Torre, 1877). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

minimus (Fabricius, 1792). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[normannus (Dejean, 1831)].Known in Denmark, Germany (Vigna Taglianti 2009), Poland (Burakowski et al. 1973).

tenellus (Erichson, 1837). Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Ferenca 2006b; Alekseev 2008c.

Trepanes Motschulsky, 1864.

articulatus (Panzer, 1796). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b 2009; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

assimilis (Gyllenhal, 1810). Ogijewicz 1933; Sharova and Grüntal 1973; Pileckis 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008 a, c; Vigna Taglianti 2009.

[clarkii (Dawson, 1849)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark, Germany (Vigna Taglianti 2009), Poland (Burakowski et al. 1973).

doris (Panzer, 1797). Jakobson 1905-1915; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

fumigatus (Duftschmid, 1812). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Alekseev 2008c.

gilvipes (Sturm, 1825). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Marggi et al. 2003; Tamutis et al. 2004; Alekseev 2008c; Vigna Taglianti 2009.

octomaculatus (Goeze, 1777). Jakobson 1905-1915; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

shuppelii (Dejean, 1831). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Alekseev 2008c.

transparens (Gebler, 1829). Monsevičius and Pankevičius 2001 (Bembidion).

Bembidion Latreille, 1802.

humerale Sturm, 1825.Jakobson 1905-1915; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

quadrimaculatum (Linnaeus, 1761). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1958b, 1960, 1976a; Pileckis et al. 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Marggi et al. 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

quadripustulatum Audinet-Serville, 1821 = quadriguttatus Olivier, 1795. # 10. Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Barševskis 2003; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

Philochthus Stephens, 1828.

aeneus (Germar, 1824). Łomnicki 1913; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Alekseev 2008c; Ivinskis et al. 2009.

biguttatus (Fabricius, 1779). Jakobson 1905-1915; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Tamutis 2000, 2005a; Marggi et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

guttula (Fabricius, 1792). Jakobson 1905-1915; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Tamutis 2000; Marggi et al. 2003; Tamutis et al. 2004, 2007; Alekseev 2008c; Vigna Taglianti 2009.

[inoptatus (Schaum, 1857)].Known in Latvia (Telnov et al. 2006).

lunulatus (Geoffroy, 1785). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

mannerheimii C.R. Sahlberg, 1827 = unicolor Chaudoir, 1850. Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bembidion); Monsevičius 1997; Marggi et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

neresheimeri P.W.J. Müller, 1929. Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Bembidion); Marggi et al. 2003; Tamutis 2005a; Alekseev 2008c; Vigna Taglianti 2009.

Ocys Stephens, 1828.

quinquestriatus (Gyllenhal, 1810). Bercio and Folwaczny 1979; Tamutis 2003; Alekseev 2008c.

Paratachys Casey, 1918.

bistriatus (Duftschmid, 1812).# 11. Pileckis and Monsevičius 1995 (Tachys); Kopecký 2003; Silfverberg 2004; Alekseev 2008c; Vigna Taglianti 2009.

micros (Fischer von Waldheim, 1828). Tamutis 2003; Barševskis 2003; Kopecký 2003; Vigna Taglianti 2009.

Porotachys Netolitzky, 1914.

bisulcatus (Nicolai, 1822). Kopecký 2003; Vigna Taglianti 2009.

Tachyura Motschulsky, 1862.

parvula (Dejean, 1831). Ferenca 2003 (Elaphropus parvulus).

Tachyta Kirby, 1837.

nana (Gyllenhal, 1810). Ogijewicz 1933; Pileckis 1960, 1976a; Jakaitis and Valenta 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Kopecký 2003; Ferenca 2003, 2004; Vaivilavičius 2008; Alekseev 2008a, c; Ivinskis et al. 1998, 2009; Vigna Taglianti 2009.

Patrobinae Kirby, 1837.

Patrobini Kirby, 1837.

Patrobus Dejean, 1821.

assimilis Chaudoir, 1844. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Alekseev 2008c; Ivinskis et al. 2009.

atrorufus Ström, 1768 = excavatus Paykull, 1790. Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Zamotailov 2003; Lynikienė 2006; Žiogas and Vaičikauskas 2007b; Alekseev 2008c; Vigna Taglianti 2009.

septentrionis Dejean, 1828 = australis J.R. Sahlberg, 1875. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Ferenca 2003; Alekseev 2008c.

Brachininae Bonelli, 1810.

Brachinini Bonelli, 1810.

Brachinus Weber, 1801.

crepitans (Linnaeus, 1758). Hrdlička 2003; Vigna Taglianti 2009.

explodens (Duftshmid, 1812). Hrdlička 2003; Vigna Taglianti 2009.

Harpalinae Bonelli, 1810.

Pterostichini Bonelli, 1810.

Stomis Clairville, 1806.

pumicatus (Panzer, 1796). Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1970a, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Bousquet 2003c; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Alekseev 2008c; Ivinskis et al. 2008, 2009; Vigna Taglianti 2009.

Poecilus Bonelli, 1810.

cupreus (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Strazdienė 1976; Dvilevičius et al. 1988; Pileckis and Šaluchaitė 1993; Silfverberg 1992, 2004; Gaidienė 1993 (Pterostichus); Pileckis and Monsevičius 1995 (Pterostichus); Monsevičius 1997, 2001; Tamutis 1999, 2000; Šablevičius 2000b; Gliaudys 2001; Bousquet 2003c; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

lepidus (Leske, 1785) = virens (O.F. Müller, 1776). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Strazdienė 1976, 1981, 1988; Silfverberg 1992, 2004; Gaidienė 1993 (Pterostichus); Pileckis and Monsevičius 1995 (Pterostichus); Monsevičius 1997; Šablevičius 2000b; Bousquet 2003c; Ferenca 2006b; Lynikienė 2006; Alekseev 2008c; Vigna Taglianti 2009.

punctulatus (Schaller, 1783). Ogijewicz 1933; Pileckis 1960, 1976a; Strazdienė 1976; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993 (Pterostichus); Pileckis and Monsevičius 1995 (Pterostichus); Tamutis 1999, 2000; Bousquet 2003c; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

versicolor (Sturm, 1824) = caerulescens auct. nec (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993 (Pterostichus); Pileckis and Monsevičius 1995 (Pterostichus); Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Gliaudys 2001; Bousquet 2003c; Tamutis et al. 2004, 2007; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Pterostichus Bonelli, 1810.

aethiops (Panzer, 1796). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Bousquet 2003c; Žiogas and Zolubas 2005; Žiogas et al. 2006; Ferenca 2006b; Lynikienė 2006 (P. atterrimus Hbst.); Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

anthracinus (Illiger, 1798). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003c; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Alekseev 2008c; Vigna Taglianti 2009.

atterimus (Herbst, 1784). Heyden 1903; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003c; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

crenatus (Duftschmid, 1812) = vernalis (Panzer, 1796) nec (O.F. Müller, 1776). Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003c; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Alekseev 2008a, c; Ivinskis et al. 2008; Vigna Taglianti 2009.

diligens (Sturm, 1824). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Ivinskis et al. 1984; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997, 2001; Bousquet 2003c; Tamutis 2000, 2005a; Tamutis et al. 2004; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Dapkus and Tamutis 2007, 2008a; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

gracilis (Dejean, 1828) = guentheri (Sturm, 1824). Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003c; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[longicollis (Duftschmid, 1812) = inaequalis (Marsham, 1802) nec (Panzer, 1796)]. Known in northern and central Poland (Burakowski et al. 1974), Denmark (Vigna Taglianti 2009).

macer (Marsham, 1802). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 1999, 2000, 2005a; Bousquet 2003c; Tamutis et al. 2006, 2007; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[madidus (Fabricius, 1775)]. Pervasive species to the European east. Known in Estonia, Poland, Denmark (Vigna Taglianti 2009).

melanarius (Illiger, 1798) = vulgaris auct. nec (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Strazdienė 1976, 1988; Dvilevičius et al. 1988; Pileckis and Šaluchaitė 1993; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 1999, 2000, 2005a; Gliaudys 2001; Bousquet 2003c; Tamutis et al. 2004, 2006, 2007; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007a, b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

minor (Gyllenhal, 1827) = brunneus (Sturm, 1824). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003c; Žiogas and Zolubas 2005; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

niger (Schaller, 1783). Heyden 1903; Raubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Strazdienė 1976, 1988; Dvilevičius et al. 1988; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Gliaudys 2001; Tamutis and Zolubas 2001; Bousquet 2003c; Tamutis et al. 2004, 2007; Žiogas et al. 2006; Ferenca 2006b; Lynikienė 2003, 2006; Žiogas and Vaičikauskas 2007b; Dapkus and Tamutis 2008a; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

nigrita (Paykull, 1790). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Strazdienė 1976, 1981, 1988; Ivinskis et al. 1984, 2008; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997, 2001; Tamutis 2000, 2005a; Bousquet 2003c; Tamutis et al. 2004; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007a, b; Alekseev 2008c; Vigna Taglianti 2009.

oblongopunctatus (Fabricius, 1787). Heyden 1903; Raubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Strazdienė 1988; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Tamutis and Zolubas 2001; Bousquet 2003c; Tamutis et al. 2004; Žiogas et al. 2006; Ferenca 2006b; Lynikienė 2003, 2006; Žiogas and Vaičikauskas 2007a, b; Dapkus and Tamutis 2007, 2008a; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

ovoideus (Sturm, 1824). Ferenca 2004, 2006a.

quadrifoveolatus Letzner, 1852 = angustatus (Duftschmid, 1812) nec (Fabricius, 1787). Barševskis 2001a; Silfverberg 2004; Alekseev 2008c; Vigna Taglianti 2009; Ivinskis et al. 2009.

rhaeticus Heer, 1837. Monsevičius and Pankevičius 2001; Barševskis 2001a; Bousquet 2003c; Silfverberg 2004; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

strenuus (Panzer, 1796). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Bousquet 2003c; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Dapkus and Tamutis 2007; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Pedius Motschulsky, 1850.

[longicollis (Duftschmid, 1812) = inaequalis (Marsham, 1802) nec (Panzer, 1796)]. Known in northern and central Poland (Burakowski et al. 1974), Denmark (Vigna Taglianti 2009).

Abax Bonelli, 1810.

[parallelepipedus (Piller & Mitterpacher, 1783)]. Known in northern Poland (Burakowski et al. 1974), Denmark, Estonia, Kaliningrad region (Vigna Taglianti 2009).

Zabrini Bonelli, 1810.

Amara Bonelli, 1810.

aenea (DeGeer, 1774). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Hieke 2003; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Tamutis et al. 2004; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[anthobia A. Villa & G.B. Villa, 1833]. Known in Latvia (Barševskis 2003), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009).

apricaria (Paykull, 1790). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Strazdienė 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

aulica (Panzer, 1796). Ogijewicz 1933; Pileckis 1960, 1976a; Strazdienė 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Hieke 2003; Ferenca 2006b; Tamutis et al. 2004, 2007 Alekseev 2008c; Vigna Taglianti 2009.

bifrons (Gyllenhal, 1810). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Strazdienė 1981, 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

brunnea (Gyllenhal, 1810). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Strazdienė 1981, 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hieke 2003; Žiogas and Zolubas 2005; Ferenca 2006b; Dapkus and Tamutis 2007; Žiogas and Vaičikauskas 2007b; Alekseev 2008c; Vigna Taglianti 2009.

communis (Panzer, 1797) = pulpani Kult, 1949 = pseudocommunis Burakowski, 1957. Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Hieke 2003; Tamutis et al. 2004; Žiogas and Zolubas 2005; Lynikienė 2006; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

concinna Zimmermann, 1832. # 12. Pileckis 1960, 1976a; Pileckis and Monsevičius 1995; Silfverberg 2004; Alekseev 2008c.

consularis (Duftschmid, 1812). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Strazdienė 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

convexior Stephens, 1828. Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hieke 2003; Alekseev 2008c; Vigna Taglianti 2009.

convexiuscula (Marsham, 1802). Hieke 2003; Vigna Taglianti 2009.

crenata (Dejean, 1828). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca 2006b; Alekseev 2008c.

cursitans Zimmermann, 1832. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Alekseev 2008c.

curta Dejean, 1828. Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

equestris (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Žiogas and Zolubas 2005; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

erratica (Duftschmid, 1812). Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Alekseev 2008c.

eurynota (Panzer, 1796). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004; Alekseev 2008c; Vigna Taglianti 2009.

famelica Zimmermann, 1832. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis 2000, 2005a; Hieke 2003; Alekseev 2008c; Vigna Taglianti 2009.

familiaris (Duftschmid, 1812). Heyden 1903; Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Hieke 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

fulva (O.F. Müller, 1776). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis and Zolubas 2001; Hieke 2003; Tamutis 2005a; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[fusca Dejean, 1828].Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark, Belarus (Vigna Taglianti 2009).

gebleri Dejean, 1831 = helleri Gredler, 1868. Barševskis 2001a; Silfverberg 2004; Ferenca et al. 2006, 2007; Alekseev 2008c; Vigna Taglianti 2009.

infima (Duftschmid, 1812). Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Alekseev 2008c; Vigna Taglianti 2009.

ingenua (Duftschmid, 1812). Miländer et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Žiogas and Zolubas 2005; Alekseev 2008c; Vigna Taglianti 2009.

littorea Thomson, 1857 = kodymi Jedlička, 1936. Monsevičius 1986a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Alekseev 2008c; Vigna Taglianti 2009.

lucida (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hieke 2003; Alekseev 2008c; Vigna Taglianti 2009.

lunicollis Schiödte, 1837 = fatrica Roubal, 1922. Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004; Žiogas and Zolubas 2005; Alekseev 2008c; Vigna Taglianti 2009.

majuscula (Chaudoir, 1850). Pileckis1970a, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hieke 2003; Tamutis 2000, 2005a; Alekseev 2008c; Vigna Taglianti 2009.

[montivaga Sturm, 1825]. Known in northern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

municipalis (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

nitida Sturm, 1825. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

ovata (Fabricius, 1792). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

plebeja (Gyllenhal, 1810). Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis 2000, 2005a; Hieke 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

praetemissa (C.R. Sahlberg, 1827). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

quenseli silvicola (Zimmermann, 1832). Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hieke 2003; Alekseev 2008c.

similata (Gyllenhal, 1810). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 1999, 2000, 2005a; Hieke 2003; Tamutis et al. 2004, 2007; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

spreta Dejean, 1831. Jakobson 1905-1915; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hieke 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[strenua Zimmermann, 1832]. Known in northern Poland (Burakowski et al. 1974), Kaliningrad region (Alekseev 2008c), Latvia (Barševskis 2003), Denmark (Vigna Taglianti 2009).

tibialis (Paykull, 1798). Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hieke 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[tricuspidata Dejean, 1831]. Known in northern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), northwestern Belarus (Alexandrovitch et al. 1996), Denmark (Vigna Taglianti 2009).

Zabrus Clairville, 1806.

tenebrioides (Goeze, 1777). Serrano and Andújar 2003; Vigna Taglianti 2009.

Sphodrini Laporte, 1834.

Calathus Bonelli, 1811.

ambiguus (Paykull, 1790). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000; Hovorka and Sciaky 2003a; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[cinctus Motschulsky, 1850]. Known in Latvia (Barševskis 2003), southern Sweden (Lundberg and Gustafsson 1995), Estonia, Poland, Denmark (Vigna Taglianti 2009).

erratus (C.R. Sahlberg, 1827) = fulvipes (Gyllenhal, 1810). Heyden 1903; Roubal, 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Strazdienė 1976, 1981; Ivinskis et al. 1984; 1988; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Žiogas and Gedminas 1994; Pileckis and Monsevičius 1995; Monsevičius 1997; Hovorka and Sciaky 2003a; Ferenca 2006b; Lynikienė 2006; Alekseev 2008c; Vigna Taglianti 2009.

fuscipes (Goeze, 1777). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1970a, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 1999, 2000, 2005a; Šablevičius 2000b; Hovorka and Sciaky 2003a; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

melanocephalus (Linnaeus, 1758). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Strazdienė 1981, 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Hovorka and Sciaky 2003a; Tamutis et al. 2004, 2007; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Alekseev 2008c; Vigna Taglianti 2009.

micropterus (Duftschmid, 1812). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Žiogas and Gedminas 1994; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hovorka and Sciaky 2003a; Žiogas and Zolubas 2005; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007a, b; Dapkus and Tamutis 2007, 2008a; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

mollis (Marsham, 1802) = ochropterus auct. nec (Duftschmid, 1812). # 13. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hovorka and Sciaky 2003a; Alekseev 2008c; Vigna Taglianti 2009.

Sphodrus Clairville, 1806

leucophtalmus (Linnaeus, 1758). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1970a, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Alekseev 2008c.

Laemostenus Bonelli, 1810 = Pristonychus Dejean, 1828.

terricola (Herbst, 1784). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ferenca 2006b; Alekseev 2008c.

Dolichus Bonelli, 1811

halensis (Schaller, 1783). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis 2000, 2005a; Barševskis 2003; Hovorka and Sciaky 2003b; Tamutis et al. 2004, 2006, 2007; Alekseev 2008c.

Synuchus Gyllenhal, 1810.

vivalis (Illiger, 1798) = nivalis (Panzer, 1797) nec (Paykull, 1790). Ogijewicz 1933; Pileckis 1960, 1976a; Strazdienė 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis 1999, 2000, 2005a; Šablevičius 2000b; Hovorka and Sciaky 2003c; Tamutis et al. 2006, 2007; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Platynini Bonelli, 1810.

Olisthopus Dejean, 1828= Odontonyx Stephens, 1827.

rotundatus (Paykull, 1790). Lentz 1879; Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1970a, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

Sericoda Kirby, 1837.

quadripunctata (DeGeer, 1774). Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agonum); Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

Anchomenus Bonelli, 1810 = Idiochroma Bedel, 1902.

dorsalis (Pontoppidan, 1763). Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Agonum); Monsevičius 1997; Tamutis 1999, 2000, 2005a; Bousquet 2003d; Tamutis et al. 2004, 2006a, 2007; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Oxypselaphus Chaudoir, 1843.

obscurus (Herbst, 1784). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Agonum); Monsevičius 1997; Gliaudys 2001; Bousquet 2003d; Žiogas and Zolubas 2005; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Paranchus Lindroth, 1974.

albipes (Fabricius, 1796) = ruficornis (Goeze, 1777) nec (DeGeer, 1774). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Agonum); Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

Platynus Bonelli, 1810.

assimilis (Paykull, 1790). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Agonum); Monsevičius 1997; Šablevičius 2000b; Bousquet 2003d; Žiogas and Zolubas 2005; Žiogas et al. 2006; Ferenca 2006b; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

krynickii (Sperk, 1835). Pileckis and Monsevičius 1982, 1995 (Agonum); Silfverberg 1992, 2004; Monsevičius 1997; Bousquet 2003d; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

livens (Gyllenhal, 1810).Pileckis 1968b, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agonum); Monsevičius 1997; Bousquet 2003d; Ferenca 2004; Žiogas and Zolubas 2005; Žiogas and Vaičikauskas 2007b; Alekseev 2008a; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

mannerheimii (Dejean, 1828). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agonum); Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

Limodromus Motschulsky, 1864.

longiventris (Mannerheim, 1825). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agonum); Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

Agonum Bonelli, 1810.

afrum (Duftschmid, 1812) = moestum auct. nec (Duftschmid, 1812). Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Barševskis 2001a; Bousquet 2003d; Ferenca 2006b; Ferenca et al. 2006, 2007; Vaivilavičius 2008; Ivinskis et al. 2008; Vigna Taglianti 2009.

[chalconotum Ménétriés, 1832 = sahlbergii (Chaudoir, 1850) = archangelicum (J.R. Sahlberg, 1875)]. Known in Latvia (Barševskis 2003).

dolens (C.R. Sahlberg, 1827). Pileckis 1960, 1976a; Sharova and Grüntal 1973; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis and Zolubas 2001; Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

duftschmidi J. Schmidt, 1994 = moestum auct. nec (Duftschmid, 1812).# 14. Barševskis 2001a; Bousquet 2003d; Silfverberg 2004; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

RDB ericeti (Panzer, 1809). Pileckis 1976a; Ivinskis et al. 1984, 2004a, 2009; Balevičius 1992; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997, 2001; Bousquet 2003d; Dapkus 2004; Tamutis 2005b; Žiogas and Zolubas 2005; Žiogas and Vaičikauskas 2005; Žiogas et al. 2006; Uselis et al. 2007; Rašomavičius 2007; Žiogas and Vaičikauskas 2007a, b; Dapkus and Tamutis 2007, 2008a, b; Vaivilavičius 2008; Alekseev 2008c, 2010b; Vigna Taglianti 2009.

fuliginosum (Panzer, 1809). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Bousquet 2003d; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Žiogas et al. 2006; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

gracile Sturm, 1824. Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003d; Žiogas et al. 2006; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

gracilipes (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003d; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

[hypocrita (Apfelbeck, 1904)].Known in northeastern Poland (Burakowski et al. 1974), Latvia (Barševskis 2003), northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

impressum (Panzer, 1796). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 1999, 2000, 2005a; Bousquet 2003d; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

lugens (Duftschmid, 1812).Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Bousquet 2003d; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

marginatum (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Strazdienė 1976; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Bousquet 2003d; Ferenca 2006b; Tamutis et al. 2008; Alekseev 2008a, c; Vigna Taglianti 2009.

micans Nicolai, 1822. Jakobson 1905-1915; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Bousquet 2003d; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

muelleri (Herbst, 1784). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Bousquet 2003d; Tamutis et al. 2004, 2007; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[munsteri (Hellén, 1935)]. Known in Latvia (Barševskis 2003), northwestern Belarus (Alexandrovitch et al. 1996), Estonia, Denmark (Vigna Taglianti 2009).

piceum (Linnaeus, 1758) = pelidnum (Herbst, 1784). Jakobson 1905-1915; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003d; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[scitulum Dejean, 1828]. Known in Poland (Burakowski et al. 1974), Latvia (Barševskis 2003), northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region, Estonia, (Vigna Taglianti 2009).

sexpunctatum (Linnaeus, 1758). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Bousquet 2003d; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

thoreyi Dejean, 1828. Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003d; Ferenca 2006b; Žiogas et al. 2006; Žiogas and Vaičikauskas 2007b; Alekseev 2008c; Ivinskis et al. 2009; Vigna Taglianti 2009.

versutum Sturm, 1824. Miländer et al. 1984; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Bousquet 2003d; Alekseev 2008c; Vigna Taglianti 2009.

viduum (Panzer, 1797). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003d; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[viridicupreum (Goeze, 1777)]. Known in Poland (Burakowski et al. 1974), Kaliningrad region, Estonia, (Vigna Taglianti 2009).

Panagaenini Bonelli, 1810.

Panagaeus Latreille, 1802.

cruxmajor (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Baehr 2003a; Inokaitis 2004; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

bipustulatus (Fabricius, 1775). Bercio and Folwaczny 1979; Ferenca 2003; Baehr 2003a; Silfverberg 2004; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Chlaeniini Brulle, 1834.

Callistus Bonelli, 1810.

[lunatus (Fabricius, 1775)]. Known in northern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

Chlaenius Bonelli, 1810.

*festivus (Panzer, 1796). # 15. Kirschenhofer 2003.

*spoliatus (P. Rossi, 1792). Pileckis and Monsevičius 1995; Barševskis 2003; disproved by Tamutis et al. (2008).

Chlaeniellus Reitter, 1908.

[kindermanni Chaudoir, 1856].Known in Latvia (Barševskis 2003), Kaliningrad region (Vigna Taglianti 2009).

nigricornis (Fabricius, 1787). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Chlaenius); Monsevičius 1997; Kirschenhofer 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

nitidulus (Schrank, 1781). Ogijewicz 1933; Pileckis 1960, 1976a; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Chlaenius); Tamutis 2000, 2005a; Ferenca 2006b; Alekseev 2008c.

tibialis Dejean, 1826. Ferenca 2006b; Vigna Taglianti 2009.

tristis (Schaller, 1783). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Chlaenius); Monsevičius 1997; Kirschenhofer 2003; Ferenca 2004, 2006b; Alekseev 2008c; Vigna Taglianti 2009.

vestitus (Paykull, 1790). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Chlaenius); Ferenca 2006b; Alekseev 2008a, c; Vigna Taglianti 2009.

Agostenus Fischer von Waldheim, 1829.

costulatus (Motschulsky, 1859). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Chlaenius); Kirschenhofer 2003; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

quadrisulcatus (Paykull, 1790). Ivinskis et al. 2008 (Chlaenius).

sulcicollis (Paykull, 1798).Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Chlaenius); Monsevičius 1997; Kirschenhofer 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

Oodini LaFerté-Sénectère, 1851.

Oodes Bonelli, 1810.

helopioides (Fabricius, 1792). Jakobson 1905-1915; Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Bousquet 2003e; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

gracilis A. Villa & G.B. Villa, 1833. Barševskis 2001a; Silfverberg 2004; Alekseev 2008c; Vigna Taglianti 2009.

Licinini Bonelli, 1810.

Badister Clairville, 1806.

bullatus (Schrank, 1798) = bipustulatus (Fabricius, 1792) nec (Fabricius, 1775). Heyden 1903; Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Baehr 2003b; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

collaris Motschulsky, 1844 = anomalus (Perris, 1866) = striatulus Hansen, 1944. Barševskis 2001; Silfverberg 2004; Vigna Taglianti 2009; Alekseev 2008c.

dilatatus Chaudoir, 1837. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Baehr 2003b; Ferenca 2004; Vaivilavičius 2008; Alekseev 2008a, c; Vigna Taglianti 2009.

dorsiger (Duftschmid, 1812).Tamutis et al. 2008; Vaivilavičius 2008.

lacertosus Sturm, 1815. Ogijewicz 1933; Pileckis 1960, 1976a; Miländer et al. 1984; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Vaivilavičius 2008; Alekseev 2008c.

meridionalis Puel, 1925. Barševskis 2001a; Silfverberg 2004; Alekseev 2008c; Vigna Taglianti 2009.

peltatus (Panzer, 1796). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c.

sodalis (Duftschmid, 1812). Sharova and Grüntal 1973; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Baehr 2003b; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

unipustulatus Bonelli, 1813. Heyden 1903; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Baehr 2003b; Žiogas and Zolubas 2005; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008 Alekseev 2008c; Vigna Taglianti 2009.

Licinus Latreille, 1802.

depressus (Paykull, 1790). Ogijewicz 1933; Pileckis 1960, 1976a; Miländer et al. 1984; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ferenca et al. 2002; Baehr 2003b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Harpalini Bonelli, 1810.

Ophonus Stephens, 1828.

[azureus (Fabricius, 1775)]. Known in Latvia (Barševskis 2003), northwestern Poland (Burakowski et al. 1974), northern Belarus (Alexandrovitch et al. 1996), southern Sweden (Lundberg and Gustafsson 1995), Denmark, Kaliningrad region (Vigna Taglianti 2009).

laticollis (Mannerheim, 1825) = nitidulus Stephens, 1828 = punctatulus (Duftschmid, 1812) nec (Fabricius, 1792). Heyden 1903; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Harpalus); Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[melletii (Heer, 1837)]. Known in northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Denmark, Kaliningrad region (Vigna Taglianti 2009).

[puncticeps Stephens, 1828]. Known in Latvia (Barševskis 2003), northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009).

puncticollis (Paykull, 1798). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Harpalus); Ferenca 2006b; Alekseev 2008c.

rufibarbis (Fabricius, 1792) = seladon (Schauberger, 1926).Heyden 1903; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Harpalus); Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

[rupicola (Sturm, 1818)].Known in Latvia (Barševskis 2003), northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Kaliningrad region (Vigna Taglianti 2009).

schaubergerianus (Puel, 1937). Tamutis et al. 2008.

[stictus Stephens, 1828]. Known in Latvia and Kaliningrad region (Vigna Taglianti 2009).

Pseudoophonus Motschulsky, 1844.

calceatus (Duftschmid, 1812). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Harpalus); Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Tamutis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

griseus (Panzer, 1796). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Harpalus); Monsevičius 1997; Kataev et al. 2003; 2004; Tamutis 2000, 2005a; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

rufipes (DeGeer, 1774) = pubescens (O.F. Müller, 1776). Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Pileckis et al. 1968; Sharova and Grüntal 1973; Pileckis and Vengeliauskaitė 1977, 1996; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Harpalus); Monsevičius 1997; Tamutis 1999, 2000, 2005a; Šablevičius 2000b; Gliaudys 2001; Kataev et al. 2003; Tamutis et al. 2004, 2006, 2007; Žiogas and Zolubas 2005; Ferenca 2006b; Lynikienė 2006; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2008c; Vigna Taglianti 2009.

Cryptophonus Brandmayr & Zetto- Brandmayr, 1882.

[*melancholicus (Dejean, 1829)]. # 16. Tamutis and Ferenca 2006; Alekseev 2008c; disproved by Tamutis et al. (2008).

Semiophonus Schauberger 1933.

signaticornis (Duftschmid, 1812). Tamutis 1999, 2000, 2005a; Silfverberg 2004; Tamutis et al. 2004; Alekseev 2008c.

Harpalus Latreille, 1802.

affinis (Schrank, 1781) = aeneus (Fabricius, 1775) nec (DeGeer, 1774). Eichwald 1830; Heyden 1903; Roubal 1910; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Pileckis and Vengeliauskaitė 1977, 1996; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 1999, 2000, 2005a; Šablevičius 2000b; Gliaudys 2001; Kataev et al. 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

anxius (Duftschmid, 1812). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kataev et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

*atratus Latreille, 1804. # 17. Pileckis 1960; Gaidienė 1993; Silfverberg 2004.

autumnalis (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

*caspius (Steven, 1806) = roubali Schauberger, 1928. # 18. Pileckis and Monsevičius 1995; Barševskis 2003.

*dimidiatus (P. Rossi, 1790). # 19. Pileckis 1960; Pileckis and Monsevičius 1995; Ferenca 2006b.

distinguendus (Duftschmid, 1812). Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis 1999, 2000, 2005a; Kataev et al. 2003; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009; Ivinskis et al. 2009.

flavescens (Piller & Mitterpacher, 1783) = rufus Brüggemann, 1873. Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kataev et al. 2003; Alekseev 2008a, c; Vigna Taglianti 2009.

froelichii Sturm, 1818. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kataev et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

hirtipes (Panzer, 1797). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Ferenca et al. 2002; Kataev et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[honestus (Duftschmid, 1812)]. Known in northwestern Poland (Burakowski et al. 1974), Belarus (Alexandrovitch et al. 1996).

laevipes Zetterstedt, 1828 = quadripunctatus Dejean, 1829. Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Kataev et al. 2003; Žiogas and Zolubas 2005; Žiogas et al. 2006; Ferenca 2006b; Lynikienė 2006; Žiogas and Vaičikauskas 2007b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

latus (Linnaeus, 1758). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Kataev et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

luteicornis (Duftschmid, 1812). Heyden 1903; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Tamutis et al. 2007; Alekseev 2008c; Vigna Taglianti 2009.

modestus (Dejean, 1829). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kataev et al. 2003; Ferenca 2006b; Vigna Taglianti 2009.

neglectus Audinet-Serville, 1821. Bercio and Folwaczny 1979; Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Alekseev 2008c.

picipennis (Duftschmid, 1812). Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Gaidienė 1993; Barševskis 2001a; Alekseev 2008c.

progrediens Schauberger, 1922. Tamutis et al. 2008.

pumilus (Sturm, 1818) = vernalis (Fabricius, 1801) nec (Panzer, 1796). Monsevičius and Pankevičius 2001; Barševskis 2001a; Silfverberg 2004; Alekseev 2008c; Vigna Taglianti 2009.

rubripes (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kataev et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

rufipalpis Sturm, 1818 = rufitarsis (Duftschmid, 1812) nec (Illiger, 1802). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

serripes (Quensel, 1806). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Barševskis 2003; Kataev et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

servus (Duftschmid, 1812). Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Kataev et al. 2003; Alekseev 2008c; Vigna Taglianti 2009.

smaragdinus (Duftschmid, 1812). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kataev et al. 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

solitaris Dejean, 1829 = fuliginosus (Duftschmid, 1812) nec (Panzer, 1809). Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Silfverberg 2004; Alekseev 2008c.

tardus (Panzer, 1796). Heyden 1903; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kataev et al. 2003; Lynikienė 2006; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

xanthopus winkleri Schauberger, 1923. Gaidienė and Ferenca 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Tamutis et al. 2008; Alekseev 2008a, c.

[zabroides Dejean, 1829]. Known in northern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

Anisodactylus Dejean, 1829.

binotatus (Fabricius, 1787). Heyden 1903; Ogijewicz 1933; Pileckis 1960, 1976a; Pileckis 1963b; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973 Silfverberg 1992, 2004; Gaidienė 1993; Šablevičius and Ferenca 1995; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Šablevičius 2000b; Tamutis and Zolubas 2001; Ito 2003; Tamutis et al. 2004, 2006, 2007; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

nemorivagus (Duftschmid, 1812). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Alekseev 2008c.

[poeciloides (Stephens, 1828)]. Known in central Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009).

[*signatus (Panzer, 1797)]. # 20. Gaidienė 1993; Silfverberg 2004; Alekseev 2008c.

Diachromus Erichson, 1837.

germanus (Linnaeus, 1758). Pileckis 1960, 1963b, 1970a, 1976a, b, 1979; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005b; Ferenca et al. 2002, 2006; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c.

Stenolophus Dejean, 1821.

[discophorus (Fischer von Waldheim, 1823)]. Known in northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009); central Poland (Burakowski et al. 1974).

mixtus (Herbst, 1784). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Acupalpus); Monsevičius 1997; Ferenca 2004; Alekseev 2008a; Ivinskis et al. 2008; Alekseev 2008c.

[skrimshiranus Stephens, 1828]. Known in northeastern Belarus (Alexandrovitch et al. 1996), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009); central Poland (Burakowski et al. 1974).

teutonus (Schrank, 1781). Ferenca 1988, 2004; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Acupalpus); Ferenca et al. 2002; Vaivilavičius 2008; Alekseev 2008c.

Bradycellus Erichson, 1837.

caucasicus (Chaudoir, 1846) = collaris (Paykull, 1798) nec (Herbst, 1784). Pileckis 1960, 1976a; Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Gaidienė 1993; Monsevičius 1997; Jaeger and Kataev 2003; Alekseev 2008c; Vigna Taglianti 2009.

[*csikii Laczó, 1912].# 21.Tamutis et al. 2008.

harpalinus (Audinet-Serville, 1821). Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2008c; Ivinskis et al. 2009.

ruficollis (Stephens, 1828) = similis (Dejean, 1829). Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004, Pileckis and Monsevičius 1995; Monsevičius 1997; Jaeger and Kataev 2003; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2008c.

[verbasci (Duftschmid, 1812)]. Known in northeastern Poland (Alexandrovitch et al. 2003), southern Sweden (Lundberg and Gustafsson 1995), Kaliningrad region (Alekseev and Bukejs 2010), Denmark (Vigna Taglianti 2009).

Dicheirotrichus Jacquelin du Val, 1857.

cognatus (Gyllenhal 1827). Pileckis and Monsevičius 1982, 1995 (Trichocellus); Silfverberg 1992, 2004; Vaivilavičius 2008; Alekseev 2008c.

[gustavii Crotch, 1871 = pubescens (Paykull, 1790) nec (O.F. Müller, 1776)]. Known in northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Vigna Taglianti 2009).

placidus (Gyllenhal 1827). Pileckis 1976a; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Trichocellus); Monsevičius 1997; Jaeger and Kataev 2003; Alekseev 2008c; Vigna Taglianti 2009.

rufithorax (C.R. Sahlberg, 1827). Pileckis 1960, 1976a, b, 1979; Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Gaidienė 1993; Jaeger and Kataev 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Acupalpus Dejean, 1829.

brunnipes (Sturm, 1825). Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Jaeger and Kataev 2003; Alekseev 2008c; Vigna Taglianti 2009.

[dubius Schilsky, 1888]. Known in Latvia (Barševskis 2003), central and northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Estonia, Denmark (Vigna Taglianti 2009).

[elegans (Dejean, 1829)]. Known in Denmark (Silfverberg 2004), Poland (Burakowski et al. 1974).

exiguus Dejean, 1829. Sharova and Grüntal 1973; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca et al. 2006; Alekseev 2008c.

flavicollis (Sturm, 1825).Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Jaeger and Kataev 2003; Alekseev 2008c; Vigna Taglianti 2009.

[interstitialis Reitter, 1884]. Known in Latvia (Barševskis 2003), Estonia (Vigna Taglianti 2009).

luteatus (Duftschmid, 1812). Ferenca et al. 2006; Tamutis and Ferenca 2006; Alekseev 2008c.

meridianus (Linnaeus, 1761). Ogijewicz 1933; Pileckis 1960, 1976a; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Tamutis and Zolubas 2001; Jaeger and Kataev 2003; Tamutis et al. 2004, 2006, 2007; Žiogas and Zolubas 2005; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

parvulus (Sturm, 1825) = dorsalis (Fabricius, 1787) nec (Pontoppidan, 1763). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Jaeger and Kataev 2003; Tamutis et al. 2004; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

suturalis Dejean, 1829. Ferenca et al. 2006; Tamutis and Ferenca 2006; Alekseev 2008c.

Anthracus Motschulsky, 1850.

consputus (Duftschmid, 1812). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Acupalpus); Monsevičius 1997; Jaeger and Kataev 2003; Alekseev 2008a, c; Ivinskis et al. 2009; Vigna Taglianti 2009.

Perigonini Horn, 1881.

Trechicus LeConte, 1853 = Perigona Laporte, 1835.

nigriceps (Dejean, 1831). Ferenca and Tamutis 2009.

Cyclosomini Laporte, 1834.

Masoreus Dejean, 1821.

wetterhallii (Gyllenhal, 1813). Mazurowa and Mazur 1939; Pileckis 1960, 1970a, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca et al. 2002, 2006; Bousquet 2003f; Žiogas and Zolubas 2005; Ivinskis and Rimšaitė 2005; Alekseev 2008c; Vigna Taglianti 2009.

Odacanthini Laporte, 1834.

Odacantha Paykull, 1798.

melanura (Linnaeus, 1767). Pileckis 1960, 1963b, 1976a; Zajančkauskas and Pileckis 1968; Sharova and Grüntal 1973; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ivinskis et al. 1998; Bousquet and Ito 2003; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

Lebiini Bonelli, 1810.

Lamprias Bonelli, 1810.

chlorocephalus (J.J. Hoffmann, 1803). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Lebia); Ferenca et al. 2002, 2006; Šablevičius 2003a, 2011; Kabak 2003; Inokaitis 2004; Vaivilavičius 2008; Alekseev 2008c; Ivinskis et al. 2009.

cyanocephala (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Lebia); Kabak 2003; Alekseev 2008c.

Lebia Latreille, 1802.

cruxminor (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kabak 2003; Alekseev 2008c; Inokaitis 2009; Vigna Taglianti 2009.

Demetrias Bonelli, 1810.

[atricapillus (Linnaeus, 1758)]. Known in Latvia (Barševskis 2003), northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Kaliningrad region (Alekseev 2008c), Denmark, Estonia (Vigna Taglianti 2009).

imperialis (Germar, 1824). Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Barševskis 2001; Ferenca 2004; Alekseev 2008c; Vigna Taglianti 2009.

monostigma Samouelle, 1819. Jakobson 1905-1915; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca et al. 2002, 2006; Šablevičius 2003a, b; Kabak 2003; Ferenca 2004; Alekseev 2008c; Vigna Taglianti 2009.

Paradromius Fowler, 1887.

linearis (Olivier, 1795). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Dromius); Barševskis 2001; Šablevičius 2001, 2003a; Ferenca et al. 2002, 2006; Alekseev 2008c; Vigna Taglianti 2009.

longiceps (Dejean, 1826). Kabak 2003; Tamutis et al. 2008; Vigna Taglianti 2009.

Dromius Bonelli, 1810.

agilis (Fabricius, 1787). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kabak 2003; Alekseev 2008c; Vigna Taglianti 2009; Ostrauskas and Ferenca 2010.

[angusticollis J.R. Sahlberg, 1880]. Known in northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Vigna Taglianti 2009).

[angustus Brullé, 1834]. Known in central and northwestern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Kaliningrad region (Alekseev 2008c), Estonia, Denmark (Vigna Taglianti 2009).

fenestratus (Fabricius, 1794). Jakobson 1905-1915; Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2001, 2003a; Alekseev 2008c.

[laeviceps Motschulsky, 1850]. Recently discovered in Latvia (Telnov et al. 2008), known in Poland (Burakowski et al. 1974), Kaliningrad region, Belarus (Vigna Taglianti 2009).

quadraticollis Morawitz, 1862. Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Alekseev 2008c; Ostrauskas and Ferenca 2010.

quadrimaculatus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kabak 2003; Ferenca 2004; Žiogas and Zolubas 2005; Vaivilavičius 2008; Alekseev 2008c; Vigna Taglianti 2009.

schneideri Crotch, 1871 = marginellus (Fabricius, 1794) nec (Herbst, 1784). Ogijewicz 1933; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Silfverberg 1992, 2004; Gaidienė 1993; Žiogas and Gedminas 1994; Pileckis and Monsevičius 1995; Kabak 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

Calodromius Reitter, 1905.

spilotus (Illiger, 1798) = quadrinotatus (Panzer, 1800) nec (Fabricius, 1798). Bercio and Folwaczny 1979; Pileckis and Monsevičius1982, 1995 (Dromius); Ferenca 1988; Silfverberg 1992, 2004; Gaidienė 1993; Alekseev 2008c.

Philorhizus Hope, 1838.

[melanocephalus (Dejean, 1825)]. Known in northern Poland (Burakowski et al. 1974), southern Sweden (Lundberg and Gustafsson 1995), Estonia, Denmark (Vigna Taglianti 2009).

notatus (Stephens, 1827) = nigriventris (Thomson, 1857). Tamutis et al. 2008; Alekseev 2008c.

[quadrisignatus (Dejean, 1825)]. Known in Belarus, Denmark, northern Russia (Vigna Taglianti 2009), southern Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1974).

sigma (P. Rossi, 1790). Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Dromius); Monsevičius 1997; Ivinskis et al. 1998; Alekseev 2008c.

Microlestes Schmidt-Goebel, 1846.

maurus (Sturm, 1827). Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Žiogas and Zolubas 2005; Ferenca 2006b; Alekseev 2008c.

minutulus (Goeze, 1777). Sharova and Grüntal 1973; Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis 2000, 2005a; Alekseev 2008c.

Syntomus Hope, 1838.

[ai Barševskis, 1993]. # 22. Known in Latvia (Barševskis 1993).

foveatus (Geoffroy, 1785). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Kabak 2003; Ferenca 2006b; Alekseev 2008c; Vigna Taglianti 2009.

[obscuroguttatus (Duftschmid, 1812)]. Known in Kaliningrad region (Bercio and Folwaczny 1979), northeastern Poland (Alexandrovitch et al. 2003).

truncatellus (Linnaeus, 1761). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Dvilevičius et al. 1988; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca et al. 2002; Kabak 2003; Alekseev 2008c; Vigna Taglianti 2009.

Lionychus Wissmann, 1846.

quadrillum (Duftschmid, 1812). Ferenca et al. 2002, 2006; Ivinskis and Rimšaitė 2005; Alekseev 2008c.

Cymindis Latreille, 1806.

angularis Gyllenhal, 1810.Pileckis 1976a; Pileckis and Monsevičius 1995; Silfverberg 1992, 2004; Ferenca 2006b; Alekseev 2008c.

*axillaris (Fabricius, 1794). Pileckis 1960, 1976a; disproved by Pileckis and Monsevičius (1995).

humeralis (Geoffroy, 1785). Miländer et al. 1984; Pileckis and Monsevičius 1995; Silfverberg 1992, 2004; Alekseev 2008c.

macularis Mannerheim 1824. Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Alekseev 2008c.

vaporariorum (Linnaeus, 1758). Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Alekseev 2008c.

HALIPLIDAE Brullé, 1835.

Brychius Thomson, 1859.

elevatus cristatus J.R. Sahlberg, 1875.Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Kovács et al. 2008; Alekseev 2010a.

Peltodytes Régimbart, 1879.

caesus (Duftschmid, 1805).Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

Haliplus Latreille, 1802.

apicalis Thomson, 1868.Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Alekseev 2010a.

confinis Stephens, 1828.Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Kovács et al. 2008; Alekseev 2010a.

flavicollis Sturm, 1834.Eichwald 1830;Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b; Ferenca 2006b; Vaivilavičius 2008; Kovács et al. 2008; Alekseev 2010a.

fluviatilis Aubé, 1836.Ogijewicz 1933; Lešinskas and Pileckis 1967; Pileckis 1960, 1976a; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

[fulvicollis Erichson, 1837]. Known in Kaliningrad region (Alekseev 2010a), northeastern Poland (Burakowski et al. 1976), Latvia (Barševskis et al. 2005), western Belarus (Alexandrovitch et al. 1996).

fulvus (Fabricius, 1801) = lapponum Thomson, 1856.Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Alekseev 2010a.

furcatus Seidlitz, 1887.Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Šablevičius 2000b; Alekseev 2010a.

heydeni Wehncke, 1875. Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Alekseev 2010a.

immaculatus Gerhardt, 1877.Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

[interjectus H. Lindberg, 1937].Known in Latvia (Barševskis et al. 2005).

laminatus (Schaller, 1783). Pileckis 1960, 1970b, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca 2006b; Alekseev 2010a.

lineatocollis (Marsham, 1802).Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca 2006b; Alekseev 2010a.

lineolatus Mannerheim, 1844 = nomax Balfour-Browne, 1911.# 23. Ogijewicz 1933; Pileckis 1960, 1976a; Pileckis and Monsevičius 1995; Monsevičius 1997.

obliquus (Fabricius, 1787).Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Kovács et al. 2008; Alekseev 2010a.

ruficollis (DeGeer, 1774).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Alekseev 2010a.

sibiricus Motschulsky, 1860 = wehnckei Gerhardt, 1877 = sahlbergi Falkenström, 1940 = lineolatus auct. nec Mannerheim, 1844. # 1.Pileckis 1976a; Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Alekseev 2010a.

variegatus Sturm, 1834. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

varius Nicolai, 1822.Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Alekseev 2010a.

NOTERIDAE Thomson, 1860.

Noterinae Thomson, 1860.

Noterini Thomson, 1860.

Noterus Clairville, 1806.

clavicornis (DeGeer, 1774). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003a, 2009; Ferenca 2006b; Alekseev 2010a.

crassicornis (O.F. Müller, 1776). Eichwald 1830; Ogijewicz 1933; Zajančkauskas and Pileckis 1968; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003a, 2009; Ferenca 2006b; Alekseev 2010a.

DYTISCIDAE Leach, 1815.

Agabinae Thomson, 1867.

Platambus Thomson, 1859.

maculatus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b, 2011; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

Agabus Leach, 1817.

affinis (Paykull, 1798). Jakobson 1905-1915; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

[*biguttulus (Thomson, 1867)]. # 24. Gaidienė 1993; Silfverberg 2004; Alekseev 2010a.

bipustulatus (Linnaeus, 1767) = solieri Aubé, 1837.Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Gaidienė 1993; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

[clypealis (Thomson, 1867)]. Known in Latvia (Barševskis et al. 2005), Kaliningrad region (Alekseev 2010a), southern Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1976), Belarus (Alexandrovitch et al. 1996).

[confinis (Gyllenhal, 1808)]. Known in Latvia (Barševskis et al. 2005).

congener (Thunberg, 1794). Ogijewicz 1933; Pileckis 1960, 1970a, 1976a, b, 1979; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

[conspersus (Marsham, 1802)].Known in northern Poland (Burakowski et al. 1976), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Nilsson 2003b, 2009).

didymus (Olivier, 1795). Gaidienė 1993; Silfverberg 2004; Alekseev 2010a.

fuscipennis (Paykull, 1798). Jakobson 1905-1915; Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

guttatus (Paykull, 1798). Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Alekseev 2010a.

labiatus (Brahm, 1791). Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

melanarius Aubé, 1837 = tarsatus Zetterstedt, 1838. Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

nebulosus (Forster, 1771). Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

paludosus (Fabricius, 1801). Lindeman 1871; Jakobson 1905-1915; Pileckis 1968b, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

[pseudoclypealis Scholtz, 1933]. Known in Latvia (Barševskis et al. 2005), Belarus (Ryndevich 2004).

[setulosus (J.R. Sahlberg, 1895)]. Known in Latvia (Barševskis et al. 2005).

striolatus (Gyllenhal, 1808). Miländer et al. 1984; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

sturmii (Gyllenhal, 1808). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

uliginosus (Linnaeus, 1761). Eichwald 1830; Lindeman 1871, Pileckis 1968b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

undulatus (Schrank, 1776). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

unguicularis (Thomson, 1867). Tamutis et al. 2008; Ivinskis et al. 2008, 2009; Alekseev 2010a.

Ilybius Erichson, 1832.

aenescens Thomson, 1870. Jakobson 1905-1915; Pileckis 1963b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Žiogas and Zolubas 2005; Ferenca 2006b; Nilsson 2009; Alekseev 2010a.

angustior (Gyllenhal, 1808). Barševskis 2001a; Silfverberg 2004; Alekseev 2010a.

ater (DeGeer, 1774). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

chalconatus (Panzer, 1796).# 13. Miländer et al. 1984 (Agabus); Gaidienė 1993 (Gaurodytes); Pileckis and Monsevičius 1982, 1995 (Agabus); Silfverberg 1992, 2004; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

crassus Thomson, 1856.Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b; Alekseev 2010a.

erichsoni (Gemminger & Harold, 1868) = nigroaeneus (Erichson, 1837) nec (Marsham, 1802). Monsevičius 1986a (Agabus); Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agabus); Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

fenestratus (Fabricius, 1781). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

fuliginosus (Fabricius, 1792). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2010a.

guttiger (Gyllenhal, 1808). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Ivinskis et al. 2008, 2009; Alekseev 2010a.

montanus (Stephens, 1828) = melanocornis (A.Zimmermann, 1915). # 13. Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

neglectus (Erichson, 1837). Monsevičius 1986a, 1998 (Agabus); Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Agabus); Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

quadriguttatus (Lacordaire, 1835) = obscurus (Marsham, 1802). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2009; Alekseev 2010a.

similis Thomson, 1856. Gaidienė 1993; Monsevičius 1997; Barševskis 2001; Silfverberg 2004; Alekseev 2010a.

subaeneus Erichson, 1837. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b; Ferenca 2006b; Alekseev 2010a.

subtilis (Erichson, 1837).Pileckis and Monsevičius 1982, 1995 (Agabus); Silfverberg 1992, 2004; Monsevičius 1997 (Agabus); Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

[*wasastjernae (C.R. Sahlberg, 1824)]. # 25. Monsevičius 1998.

Colymbetinae Erichson, 1837.

Colymbetini Erichson, 1837.

Rhantus Dejean, 1833.

bistriatus (Bergsträsser, 1778) = adspersus (Fabricius, 1801).Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

consputus (Sturm, 1834). Pileckis and Monsevičius 1995; Alekseev 2010a.

exsoletus (Forster, 1771). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2003; Nilsson 2003b, 2009; Ivinskis et al. 2008; Alekseev 2010a.

frontalis (Marsham, 1802) = notatus (Fabricius, 1781) nec (Bergsträsser, 1778). Lentz 1879; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

grapii (Gyllenhal, 1808).Jakobson 1905-1915; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

incognitus (R. Scholz, 1927). Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Alekseev 2010a.

latitans Sharp, 1882. Barševskis 2001a; Silfverberg 2004; Alekseev 2010a.

notaticollis (Aubé, 1837). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

suturalis (MacLeay, 1825) = pulverosus (Stephens, 1828).Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

suturellus (Harris, 1828). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2003a; Nilsson 2003b, 2009; Alekseev 2010a.

Colymbetes Clairville, 1806.

fuscus (Linnaeus, 1758). Eichwald 1830; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

paykulli Erichson, 1837. Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

striatus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

Copelatinae Van den Branden, 1885.

Copelatini Van den Branden, 1885.

Copelatus Erichson, 1832.

haemorrhoidalis (Fabricius, 1787) = ruficollis (Schaller, 1783) nec (DeGeer, 1774). Pileckis 1960, 1976a; Pileckis and Monsevičius 1995; Silfverberg 1992, 2004; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

Dytiscinae Leach, 1815.

Hydaticini Sharp, 1880.

Hydaticus Leach, 1817.

[aruspex Clark, 1864 = laevipennis Thomson, 1867]. Known in Latvia (Barševskis et al. 2005), Kaliningrad region (Alekseev 2010a), southern Sweden (Lundberg and Gustafsson 1995), northern Poland (Burakowski et al. 1976), Belarus (Alexandrovitch et al. 1996).

continentalis Balfour-Browne, 1944 = stagnalis (Fabricius, 1787) nec (Geoffroy, 1785).Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2003a; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

seminiger (DeGeer, 1774). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

transversalis (Pontoppidan, 1763). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

Aciliini Thomson, 1867.

Graphoderus Dejean, 1833.

austriacus (Sturm, 1834).Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Alekseev 2010a.

RDB bilineatus (DeGeer, 1774). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Ferenca 2004, 2006a; Tamutis 2005b; Kriaučiūnienė and Zaplatkin 2007; Rašomavičius 2007; Uselis et al. 2007; Ivinskis et al. 2008, 2009; Švitra and Aliukonis 2009; Alekseev 2010a, b; Šablevičius 2011.

cinereus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2009; Alekseev 2010a.

zonatus zonatus (Hoppe, 1795). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

Acilius Leach, 1817.

canaliculatus (Nicolai, 1822). Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2010a; Šablevičius 2011.

sulcatus (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

Dytiscini Leach, 1815.

Dytiscus Linnaeus, 1758.

circumcinctus Ahrens, 1811. Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

circumflexus Fabricius, 1801. Pileckis 1960, 1970a, b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

dimidiatus Bergsträsser, 1778. Ogijewicz 1933; Pileckis 1960, 1970a, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

lapponicus Gyllenhal, 1808. Gaidienė and Ferenca 1988; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000a, 2003a, b, 2011; Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

RDB latissimus Linnaeus, 1758. Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Gliaudys 2001; Nilsson 2003b, 2009; Tamutis 2005b; Ferenca 2006b; Rašomavičius 2007; Lopeta 2007a; Ivinskis et al. 2008; Aliukonis and Švitra 2009; Švitra and Aliukonis 2009; Alekseev 2010a, b.

marginalis Linnaeus, 1758 = conformis Kunze, 1818.Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b, 2011; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

semisulcatus O.F. Müller, 1776 = punctulatus Fabricius, 1776. Ogijewicz 1993; Pileckis 1960, 1970a; Silfverberg 1992, 2004; 1976a; Gaidienė 1993; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

Cybisterini Sharp, 1880.

Cybister Curtis, 1827.

lateramarginalis (DeGeer, 1774). Ogijewicz 1933; Pileckis 1960, 1970a, b, 1976a; Lešinskas and Pileckis 1967; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Gliaudys 2001; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a; Šablevičius 2011.

Hydroporinae Aubé, 1836.

Laccornini Wolfe & Roughley, 1990.

Laccornis Gozis, 1914.

oblongus (Stephens, 1835).# 26. Monsevičius 1997.

Hydrovatini Sharp, 1880.

Hydrovatus Motschulsky, 1853.

[cuspidatus (Kunze, 1818)]. Known in northern Poland (Burakowski et al. 1976), Denmark (Nilsson 2003b, 2009).

Bidessini Sharp, 1880.

Bidessus Sharp, 1880.

grossepunctatus Vorbringer, 1907.Barševskis 2001a; Silfverberg 2004; Alekseev 2010a.

unistriatus (Goeze, 1777).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

Hydroglyphus Motschulsky, 1853 = Guignotus Houlbert, 1934.

geminus (Fabricius, 1792) = pusillus (Fabricius, 1781) nec (O.F. Müller, 1776). Miländer et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995 (Bidessus); Šablevičius 2003a. Nilsson 2003b, 2009; Alekseev 2010a.

hamulatus (Gyllenhal, 1813). Pileckis and Monsevičius 1995 (Bidessus); Silfverberg 2004; Alekseev 2010a.

Hyphydrini Gistel, 1848.

Hyphydrus Illiger, 1802.

ovatus (Linnaeus, 1761).Ogijewicz 1933; Pileckis 1960, 1976a; Ivinskis et al. 1984; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a; Šablevičius 2011.

Hygrotini Portevin, 1929.

Hygrotus Stephens, 1828.

[confluens (Fabricius, 1787)].Known in Kaliningrad region (Alekseev 2010a), northeastern Poland (Burakowski et al. 1976), eastern Belarus (Alexandrovitch et al. 1996); southern Sweden (Lundberg and Gustafsson 1995), Denmark (Nilsson 2003b, 2009).

decoratus (Gyllenhal, 1810).Jakobson 1905-1915; Monsevičius 1986a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2003a; Nilsson 2003b, 2009; Alekseev 2010a.

impressopunctatus (Schaller, 1783).Ogijewicz 1933 (Coelambus); Pileckis 1960, 1976a (Coelambus); Silfverberg 1992, 2004 (Coelambus);Gaidienė 1993 (Coelambus); Pileckis and Monsevičius 1995 (Coelambus); Monsevičius 1997 (Coelambus); Nilsson 2003b, 2009; Alekseev 2010a.

inaequalis (Fabricius, 1776).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

[marklini (Gyllenhal, 1813)].Known in Latvia (Barševskis et al. 2005); Kaliningrad region (Alekseev 2010a), northwestern Belarus (Alexandrovitch et al. 1996), northern Poland (Burakowski et al. 1976).

[nigrolineatus (Steven, 1808) = lautus (Schaum, 1843)].Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Nilsson 2003b, 2009), Belarus (Ryndevich 2004).

novemlineatus (Stephens, 1829). Tamutis et al. 2008 (Coelambus); Alekseev 2010a.

parallelogrammus (Ahrens, 1812). Karalius and Monsevičius 1992 (Coelambus); Pileckis and Monsevičius 1995 (Coelambus); Silfverberg 1996, 2004 (Coelambus); Alekseev 2010a.

polonicus (Aubé, 1842). Silfverberg 1992, 2004 (Coelambus); Gaidienė 1993 (Coelambus); Pileckis and Monsevičius 1995 (Coelambus); Ivinskis et al. 2009; Alekseev 2010a.

[quinquelineatus (Zetterstedt, 1828)]. Known in northern Belarus (Alexandrovitch et al. 1996), southern Sweden (Lundberg and Gustafsson 1995), Denmark (Nilsson 2003b, 2009).

versicolor (Schaller, 1783). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Vaivilavičius 2008; Alekseev 2010a.

Hydroporini Aubé, 1836.

Hydroporus Clairville, 1806 = Hydrocolus Roughley & Nilsson, 2000.

angustatus Sturm, 1835. Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

[brevis R.F. Sahlberg, 1834]. Known in Kaliningrad region (Alekseev 2010a), northeastern Poland (Burakowski et al. 1976), northwestern Belarus (Alexandrovitch et al. 1996), southern Sweden (Lundberg and Gustafsson 1995), Latvia (Barševskis et al. 2005).

elongatulus Sturm, 1835 = eugeniae Zaitzev, 1909.Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

erythrocephalus (Linnaeus, 1758). Lentz 1879; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

fuscipennis Schaum, 1868. Pileckis 1960, 1970a, 1976a, b, 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b Alekseev 2010a.

glabriusculus Aubé, 1838.Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

[gyllenhalii Schiödte, 1841 = piceus auct. nec Stephens, 1828].Known in northeastern Poland (Burakowski et al. 1976), southern Sweden (Lundberg and Gustafsson 1995), Belarus, Denmark (Nilsson 2003b, 2009).

incognitus Sharp, 1869. Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Monsevičius 1997; Alekseev 2010a.

[kraatzii Schaum, 1867]. Known in Poland (Burakowski et al. 1976), Estonia (Nilsson 2003b, 2009).

longicornis Sharp, 1871. Miländer et al. 1984; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

melanarius Sturm, 1835. Lentz 1879; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

memnonius Nicolai, 1822. Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2006b; Nilsson 2003b, 2009; Alekseev 2010a.

morio Aubé, 1838 = valliger Hellén, 1929 = melanocephalus auct. nec (Marsham, 1802).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

neglectus Schaum, 1845. Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

[neuter Fairmaire & Laboulbène, 1854 = discretus Fairmaire, 1859].Known in Kaliningrad region (Alekseev 2010a), northern Poland (Burakowski et al. 1976), Latvia (Barševskis et al. 2005), southern Sweden (Lundberg and Gustafsson 1995), northwestern Belarus (Alexandrovitch et al. 1996).

[nigellus Mannerheim, 1853 = tartaricus LeConte, 1850].Known in Latvia (Barševskis et al. 2005), southern Sweden (Lundberg and Gustafsson 1995).

nigrita (Fabricius, 1792). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

notatus Sturm, 1835.Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Ferenca et al. 2002; Alekseev 2010a.

obscurus Sturm, 1835.Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

[obsoletus Aubé, 1838]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Nilsson 2003b, 2009).

palustris (Linnaeus, 1761). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Ferenca 2006b; Nilsson 2003b, 2009; Alekseev 2010a.

planus (Fabricius, 1781). Silfverberg 1992, 2004; Gaidienė 1993; Šablevičius and Ferenca 1995; Pileckis and Monsevičius 1995; Monsevičius 1997; Vaivilavičius 2008; Nilsson 2003b, 2009; Ivinskis et al. 2009; Alekseev 2010a.

pubescens (Gyllenhal, 1808). Tamutis 2003; Alekseev 2010a.

rufifrons (O.F. Müller, 1776) = piceus Stephens, 1828.# 27. Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

scalesianus Stephens, 1828. Ferenca and Tamutis 2009; Alekseev 2010a.

striola (Gyllenhal, 1826). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ivinskis et al. 2008; Alekseev 2010a.

[tessellatus (Drapiez, 1819)]. Known in Latvia (Barševskis et al. 2005).

tristis (Paykull, 1798). Ogijewicz 1933; Pileckis 1960, 1970a, 1976a, b, 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

umbrosus (Gyllenhal, 1808). Jakobson 1905-1915; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

Porhydrus Guignot, 1945.

lineatus (Fabricius, 1775). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Alekseev 2010a.

[obliquesignatus (Bielz, 1852)]. Known in northwestern Belarus (Alexandrovitch et al. 1996).

Graptodytes Seidlitz, 1887.

bilineatus (Sturm, 1835). Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Šablevičius 2000b; Alekseev 2010a.

granularis (Linnaeus, 1767). Jakobson 1905-1915; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

pictus (Fabricius, 1787). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2003a; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

Oreodytes Seidlitz, 1887.

sanmarkii (C.R. Sahlberg, 1826) = rivalis (Gyllenhal, 1827). # 28. Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

Suphrodytes DesGozis, 1914.

dorsalis (Fabricius, 1787) = eljasi (Wirén, 1968).Ogijewicz 1933 (Hydroporus); Pileckis 1960, 1976a (Hydroporus); Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Hydroporus); Monsevičius 1997 (Hydroporus); Šablevičius 2000b, 2003b (Hydroporus); Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

Deronectes Sharp, 1880.

latus (Stephens, 1829). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

Scarodytes DesGozis, 1914.

halensis (Fabricius, 1787). Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Gaidienė 1993; Monsevičius 1997; Šablevičius 2000b, 2011; Nilsson 2003b, 2009; Alekseev 2010a.

Nebrioporus Régimbart, 1906 = Potomonectes A.Zimmermann, 1921.

assimilis (Paykull, 1798). Pileckis and Monsevičius 1995; Silfverberg 2004; Alekseev 2010a.

canaliculatus (Lacordaire, 1835).Nilsson 2003b.

depressus (Fabricius, 1775) = elegans (Panzer, 1794). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Nilsson 2003b, 2009; Alekseev 2010a.

Stictotarsus A. Zimmermann, 1919.

duodecimpustulatus (Fabricius, 1792). # 29. Pileckis 1960 (Deronectes); Ferenca 2006b.

[griseostriatus (DeGeer, 1774)]. Known in souhern Sweden, Estonia (Lundberg and Gustafsson 1995).

Laccophilinae Gistel, 1848.

Laccophilini Gistel, 1848.

Laccophilus Leach, 1815.

hyalinus (DeGeer, 1774).Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

minutus (Linnaeus, 1758) = obscurus (Panzer, 1794). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

poecilus Klug, 1834 = ponticus Sharp, 1882 = variegatus (Germar, 1812) nec (Geoffroy, 1785) = obsoletus auct. nec Westhoff, 1881. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Gliaudys 2001; Nilsson 2003b, 2009; Ferenca 2006b; Alekseev 2010a.

POLYPHAGA Emery, 1886.

HYDROPHILOIDEA Latreille, 1802.

HYDROPHILIDAE Latreille, 1802.

Helophorinae Leach, 1815.

Helophorus Fabricius, 1775.(Hydraenidae)

[aequalis Thomson, 1868]. Known in Poland, Denmark (Hansen 2009), southern Sweden (Lundberg and Gustafsson 1995), Latvia (Telnov 2004).

aquaticus (Linnaeus, 1758). Eichwald 1830; Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Löbl and Smetana 2004; Ferenca 2006b; Vaivilavičius 2008; Hansen 2004, 2009; Alekseev 2010a.

arvenicus Mulsant, 1846. Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004a, 2009; Vorst et al. 2007; Alekseev 2010a.

[asperatus Rey, 1885]. Known in southern Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1976), Denmark (Hansen 2009).

brevipalpis Bedel, 1881. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004a, 2009; Ferenca 2006b; Alekseev 2010a.

[croaticus Kuwert, 1886]. Known in Belarus (Ryndevich 2004), Poland (Burakowski et al. 1976), recently discovered in Latvia (Telnov et al. 2006).

flavipes Fabricius, 1792 = viridicollis Stephens, 1828. Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004a, 2009; Ferenca 2006b; Alekseev 2010a.

[fulgidicollis Motschulsky, 1860]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Hansen 2009).

grandis Illiger, 1798. Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Hansen 2004a; Hansen 2009; Alekseev 2010a.

granularis (Linnaeus, 1761). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1968b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004a, 2009; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2010a.

griseus Herbst, 1793 = affinis Marsham, 1802. Pileckis 1968b, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Šablevičius and Ferenca 1995; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004a, 2009; Ferenca 2006b; Vaivilavičius 2008; Ivinskis et al. 2009; Alekseev 2010a.

[lapponicus Thomson, 1853]. Known in northwestern Belarus (Alexandrovitch et al. 1996; Ryndevich 2004), eastern Prusia (Schilsky 1909), Sweden (Lundberg and Gustafsson 1995), Estonia, Denmark (Hansen 2009).

[laticollis Thomson, 1853]. Known in Kaliningrad region (Alekseev 2010a), Belarus (Ryndevich 2004), northwestern Poland (Burakowski et al. 1976), Sweden (Lundberg and Gustafsson 1995), Estonia, Denmark (Hansen 2009).

longitarsis Wollaston, 1864. Monsevičius 1997.

minutus Fabricius, 1775. Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004a, 2009; Alekseev 2010a.

nanus Sturm, 1836. Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004a, 2009; Alekseev 2010a.

nubilus Fabricius, 1777. Eichwald 1830; Lindeman 1871; Pileckis 1968b; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004a, 2009; Alekseev 2010a.

[obscurus Mulsant, 1844 = walkeri Sharp, 1916]. Known in southern Sweden (Lundberg and Gustafsson 1995), northern Poland (Burakowski et al. 1976), Denmark (Hansen 2009).

[paraminutus Angus, 1986].Known in northwestern Belarus (Alexandrovitch et al. 1996; Ryndevich 2004).

[*pumilio Erichson, 1837]. # 30. Lindeman 1871; Pileckis 1968b (Helophorus pumilio Er. = Helophorus redtenbacheri Kuv., 1976a (Helophorus pumilio Er. = Helophorus redtenbacheri Kunc.); Alekseev 2010a.

[*redtenbacheri Kuwert, 1885]. # 30. Silfverberg 1992, 2004; Pileckis and Monsevičius 1995 (Helophorus redtenbacheri Kuwert = Helophorus pumilio auct. nec Er.); Hansen 2004a, 2009; Alekseev 2010a.

strigifrons Thomson, 1868. Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Alekseev 2010a.

tuberculatus Gyllenhal, 1810. Bercio and Folwaczny 1979; Monsevičius 1986a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Ferenca 2004; Hansen 2004a, 2009; Alekseev 2010a.

Georissinae Laporte, 1840.

Georissus Latreille, 1809.(Georissidae)

crenulatus (P. Rossi, 1794). Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Šablevičius 2000a, 2003a; Hansen 2004b, 2009.

Hydrochinae Thomson, 1859.

Hydrochus Leach, 1817.(Hydraenidae)

[angustatus Germar, 1824]. Known in Latvia (Telnov 2004), central Poland (Burakowski et al. 1976), Estonia (Silfverberg 1992, 2004).

brevis (Herbst, 1793). Lindeman 1871; Pileckis 1968b, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Barševskis 2001a; Hansen 2004c; Vaivilavičius 2008; Alekseev 2010a.

crenatus (Fabricius, 1792) = carinatus Germar, 1824. Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Barševskis 2001a; Ferenca et al. 2006; Alekseev 2010a.

elongatus (Schaller, 1783). # 31. Ogijewicz 1933; Pileckis 1960, 1976a; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Ivinskis et al. 2008.

ignicollis Motschulsky, 1860. # 31. Silfverberg 1992, 2004; Hansen 2004c, 2009; Ferenca 2006b; Alekseev 2010a.

[kirgisicus Motschulsky, 1860]. Known in central Russia (Shatrovskij 1993), northwestern Belarus (Alexandrovitch et al. 1996; Ryndevich 2004).

[megaphallus van Berge Henegouwen, 1988]. Recently found in Latvia (Telnov et al. 2006), known in Belarus (Ryndevich 2004), Sweden (Lundberg and Gustafsson 1995), Poland, Denmark (Hansen 2009).

Spercheinae Erichson, 1837.

Spercheus Illiger, 1798.(Sphercheidae)

emarginatus (Schaller, 1783). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Šablevičius 2000b, 2003a; Hansen 2004d, 2009; Alekseev 2010a.

Hydrophilinae Latreille, 1802.

Berosini Mulsant, 1844

Berosus Leach, 1817.

[fulvus Kuwert, 1888]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Hansen 2009).

luridus (Linnaeus, 1761). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

signaticollis (Charpentier, 1825). Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Alekseev 2010a.

[spinosus (Steven, 1808)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Latvia, Estonia (Hansen 2009), central Poland (Burakowski et al. 1976), Belarus (Ryndevich 2004).

Chaetarthriini Bedel, 1881

Chaetarthria Stephens, 1832.

seminulum (Herbst, 1797). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Ivinskis et al. 2008; Šablevičius 2011.

Anacaenini Hansen, 1991

Paracymus Thomson, 1867.

[aeneus (Germar, 1824)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Hansen 2009).

Anacaena Thomson, 1859.

globulus (Paykull, 1798).Ivinskis et al. 2008, 2009; Alekseev 2010a.

lutescens (Stephens, 1832) = limbata auct. nec (Fabricius, 1792). # 32. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis and Zolubas 2001; Hansen 2004e, 2009; Vaivilavičius 2008; Alekseev 2010a.

limbata (Fabricius, 1792).# 32. Ferenca and Tamutis 2009; Alekseev 2010a.

Laccobiini Houlbert, 1922

Laccobius Erichson, 1837.

[albipes Kuwert, 1890]. Known in northern Poland (Burakowski et al. 1976), recently found in Latvia (Vorst et al. 2007).

bipunctatus (Fabricius, 1775). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

[colon (Stephens, 1829) = biguttatus Gerhardt, 1877]. Known in Kaliningrad region (Alekseev 2010a), southern Sweden (Lundberg and Gustafsson 1995), northern Poland (Burakowski et al. 1976), Latvia (Telnov 2004), northwestern Belarus (Alexandrovitch et al. 1996), Denmark (Hansen 2009).

[decorus (Gyllenhal, 1827)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Estonia (Hansen 2009), recently found in Latvia (Vorst et al. 2007).

minutus (Linnaeus, 1758).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius and Ferenca 1995; Šablevičius 2000b, 2011; Hansen 2004e, 2009; Ferenca 2006b; Vaivilavičius 2008; Alekseev 2010a.

sinuatus Motschulsky, 1849. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Hansen 2004e, 2009; Alekseev 2010a.

striatulus (Fabricius, 1801). Karalius and Monsevičius 1992; Pileckis and Monsevičius 1995; Silfverberg 1996, 2004; Alekseev 2010a.

Hydrophilini Latreille, 1802.

Helochares Mulsant, 1844.

obscurus (O.F. Müller, 1776) = erythrocephalus (Fabrius 1792). # 33. Ogijewicz 1933; Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

*lividus (Forster, 1771) = griseus (Fabricius, 1787). # 33. Pileckis 1976a; Silfverberg 1992, 2004; Hansen 2004e; Pileckis and Monsevičius 1995.

Enochrus Thomson, 1859.

affinis (Thunberg, 1794). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

bicolor (Fabricius, 1792). Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ferenca 2006a; Alekseev 2010a.

coarctatus (Gredler, 1863). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

fuscipennis (Thomson, 1884) = sahlbergi (Fauvel, 1887). # 34. Pileckis 1960; Pileckis 1976a; Gaidienė 1993; Pileckis and Monsevičius 1995; Silfverberg 2004; Ferenca 2006b; Alekseev 2010a.

[halophilus (Bedel, 1878)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Denmark (Hansen 2009).

melanocephalus (Olivier, 1792). Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ivinskis et al. 2009; Alekseev 2010a.

ochropterus (Marsham, 1802) = frontalis (Erichson, 1837). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004e, 2009; Alekseev 2010a.

quadripunctus (Herbst, 1797). # 34.Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

testaceus (Fabricius, 1801). Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

Cymbiodyta Bedel, 1881.

marginella (Fabricius, 1792).Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Gaidienė 1993; Monsevičius 1997; Hansen 2004e, 2009; Vaivilavičius 2008; Alekseev 2010a.

Hydrobius Leach, 1815.

fuscipes (Linnaeus, 1758) = subrotundus Stephens, 1829 = rottenbergi Gerhardt, 1872. Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Ferenca 2006b; Ivinskis et al. 2008; Hansen 2004e, 2009; Alekseev 2010a.

Limnoxenus Motschulsky, 1853.

niger (Gmelin, 1790) = picipes (Fabricius, 1787). Pileckis and Monsevičius 1995.

Hydrochara Berthold, 1827.

caraboides (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1963b, 1976a; Lešinskas and Pileckis 1967; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b, 2011; Gliaudys 2001; Hansen 2004e, 2009; Ferenca 2006b; Alekseev 2010a.

Hydrophilus Geoffroy, 1762 = Hydrous Linnaeus, 1775.

aterrimus Eschscholtz, 1822. Ogijewicz 1933; Pileckis 1960, 1963, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a; Šablevičius 2011.

piceus (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Pileckis 1960, 1970a, b, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Gliaudys 2001; Hansen 2004e, 2009; Ferenca 2006b; Ivinskis et al. 2008; Alekseev 2010a.

Sphaeridiinae Latreille, 1802.

Coelostomatini Heyden, 1891 (1890)

Coelostoma Brullé, 1835.

orbiculare (Fabricius, 1775).Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ferenca 2006b; Monsevičius 1997; Vaivilavičius 2008; Ivinskis et al. 2008; Alekseev 2010a; Šablevičius 2011.

Megasternini Mulsant, 1844.

Cercyon Leach, 1817.

analis (Paykull, 1798). Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Šablevičius 2011.

bifenestratus Küster, 1851. Ogijewicz 1933; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ivinskis et al. 2009.

castaneipenis Vorst, 2009. Known in Latvia, Poland, Sweden (Vorst 2009).

convexiusculus Stephens, 1829. Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b.

[depressus Stephens, 1829]. Known in southern Sweden (Lundberg and Gustafsson 1995), Estonia, Denmark (Hansen 2009).

granarius Erichson, 1837. Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b.

haemorrhoidalis (Fabricius, 1775). Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Tamutis and Zolubas 2001; Hansen 2004e, 2009; Ferenca 2006b.

impressus (Sturm, 1807). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Žiogas and Zolubas 2005.

laminatus Sharp, 1873. Monsevičius 1986a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ivinskis et al. 2009.

lateralis (Marsham, 1802). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Ryndevich 2004; Hansen 2004e, 2009; Ferenca 2006b.

littoralis (Gyllenhal, 1808). Pileckis 1960, 1976a, b, 1979; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ferenca 2006b.

marinus Thomson, 1853. Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004e, 2009; Ivinskis et al. 2009.

melanocephalus (Linnaeus, 1758). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Hansen 2004e, 2009; Ferenca 2006b.

nigriceps (Marsham, 1802) = atricapillus (Marsham, 1802). Tamutis et al. 2008.

obsoletus (Gyllenhal, 1808) = lugubris auct. nec(Olivier, 1790). Gaidienė 1993; Silfverberg 2004.

pygmaeus (Illiger, 1801). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b; Šablevičius 2011.

quisquilius (Linnaeus, 1761). Ogijewicz 1933; Pileckis 1960, 1976a; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b.

sternalis Sharp, 1928. Tamutis et al. 2008; Ivinskis et al. 2008.

terminatus (Marsham, 1802). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Hansen 2004e, 2009.

tristis (Illiger, 1801). Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Ferenca 2006b.

unipunctatus (Linnaeus, 1758).Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009; Šablevičius 2011.

ustulatus (Preyssler, 1790). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000b; Hansen 2004e, 2009; Ferenca 2006b.

Megasternum Mulsant, 1844.

concinnum (Marsham, 1802) = obscurum (Marsham, 1802) = boletophagum (Marsham, 1802) nec Marsham, 1802. Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Hansen 2004e, 2009.

Cryptopleurum Mulsant, 1844.

crenatum (Kugelann, 1794). Monsevičius 1986a, 1997; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004e, 2009.

minutum (Fabricius, 1775). Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004e, 2009; Ferenca 2006b.

[subtile Sharp, 1884]. Known in southern Sweden (Lundberg and Gustafsson 1995), Estonia (Süda 2009), northwestern Belarus (Alexandrovitch et al. 1996), Denmark (Hansen 2009).

Sphaeridini Latreille, 1802.

Sphaeridium Fabricius, 1775.

bipustulatum Fabricius, 1781. Eichwald 1830; Roubal 1910; Ogijewicz 1933; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b; Hansen 2004e, 2009; Ferenca 2006b.

lunatum Fabricius, 1792. Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Monsevičius 1997; Tamutis and Zolubas 2001; Hansen 2004e, 2009.

marginatum Fabricius, 1787. Monsevičius and Pankevičius 2001; Silfverberg 2004.

scarabaeoides (Linnaeus, 1758). Eichwald 1830; Ogijewicz 1933; Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Lešinskas and Pileckis 1967; Zajančkauskas and Pileckis 1968; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2011; Hansen 2004e, 2009; Ferenca 2006b.

substriatum Faldermann, 1838. Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Hansen 2004e, 2009.

SPHAERITIDAE Shuckard, 1839.

Sphaerites Duftschmid, 1805.

glabratus (Fabricius, 1792). Pileckis 1968a, 1970a, 1976a, b, 1979; Pileckis and Monsevičius 1995; Silfverberg 1992, 2004; Ferenca 2004; Löbl 2004a, 2009.

HISTERIDAE Gyllenhal, 1808.

Abraeinae MacLeay, 1819.

Abraeini MacLeay, 1819.

Chaetabraeus Portevin, 1929.

globulus (Creutzer, 1799). Mazurowa and Mazur 1939 (Abraeus); Pileckis 1960, 1976a (Abraeus); Pileckis and Monsevičius 1995; Silfverberg 2004; Mazur 2004.

Abraeus Leach, 1817.

[granulum Erichson, 1839]. Known in southern Sweden (Lundberg and Gustafsson 1995), northwestern Belarus (Alexandrovitch et al. 1996), Kaliningrad region (Alekseev and Bukejs 2010), northern Poland (Burakowski et al. 1978; Mazur 2004); Denmark (Lackner 2009).

[parvulus Aubé, 1842]. Known in southern Sweden (Lundberg and Gustafsson 1995), northeastern Poland (Burakowski et al. 1978; Mazur 2004), Denmark (Lackner 2009).

perpusillus (Marsham, 1802) = globosus (Hoffmann, 1803). Pileckis and Monsevičius 1995; Silfverberg 2004; Mazur 2004.

Plegaderini Portevin, 1929.

Plegaderus Erichson, 1834.

caesus (Herbst, 1792). Pileckis 1968b; Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Mazur 2004; Lackner 2009.

dissectus Erichson, 1839. # 35. Lentz 1879; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995.

saucius Erichson, 1834. Pileckis 1976a; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Šablevičius 2000a, 2001, 2003a, b, 2011; Mazur 2004; Lackner 2009.

vulneratus (Panzer, 1797). Jakaitis 1973; Pileckis 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Šablevičius 2000b, 2003a, b; Mazur 2004; Lackner 2009.

Acritini Wenzel, 1944.

Acritus LeConte, 1853.

[homoeopathicus Wollaston, 1857]. Known in Latvia (Telnov 2004), Poland (Burakowski et al. 1978; Mazur 2004); Denmark (Lackner 2009).

minutus (Herbst, 1792). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Mazur 2004; Lackner 2009.

[nigricornis (Hoffmann, 1803)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Latvia (Telnov 2004), northern Poland (Burakowski et al. 1978; Mazur 2004); Denmark (Lackner 2009).

Aeletes Horn, 1873.

[atomarius (Aubé, 1842)]. Known in southern Sweden (Lundberg and Gustafsson 1995), Poland (Burakowski et al. 1978; Mazur 2004); Denmark (Lackner 2009).

Teretriini Bickhardt, 1914.

Teretrius Erichson, 1834.

fabricii Mazur, 1972 = picipes (Fabricius, 1787) nec (Olivier, 1789). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Tamutis and Zolubas 2001; Mazur 2004; Lackner 2009.

Saprininae Blanchard, 1845.

Saprinus Erichson, 1834.

aeneus (Fabricius, 1775). Mazurowa and Mazur 1939; Pileckis 1960, 1976a; Bercio and Folwaczny 1979; Silfverberg 1992, 2004; Gaidienė 1993; Pileckis and Monsevičius 1995; Monsevičius 1997; Mazur 2004; Lackner 2009.

immundus (Gyllenhal, 1827). Pileckis 1976a; Silfverberg 1992, 2004; Pileckis and Monsevičius 1995; Mazur 2004; Ferenca et al. 2006; Lackner 2009.

planiusculus Motschulsky, 1849 = cuspidatus Ihssen, 1949. Pileckis and Monsevičius 1982, 1995; Silfverberg 1992, 2004; Mazur 2004; Ferenca et al. 2006.

rugifer (Paykull, 1809) = quadristriatus auct. nec (Thunberg, 1794). Tenenbaum 1931; Pileckis